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fineleaf fournerve daisy, slimleaf actinea

perky sue, stemless four-nerve daisy, stemless hymenoxys, stemless woollybase

Habit Annuals, 16–50+ cm. Perennials, 2–30+ cm.
Caudices

none.

± branched, branches notably thickened distally.

Stems

1–10, erect or ± decumbent, leafy, unbranched or branched distally.

1–35(–60), erect, unbranched.

Leaves

basal and cauline, ± hairy, ± gland-dotted;

basal blades spatulate to oblanceolate, entire or with 2–6 teeth or lobes;

proximal cauline blades narrowly spatulate or oblanceolate to linear-oblanceolate, entire or with 2–6 teeth or lobes;

mid blades oblanceolate to linear-oblanceolate to linear, 0.9–9.5(–16) mm wide, entire or with 1–2 teeth or lobes;

distal blades narrowly oblanceolate to linear-oblanceolate to linear, entire or with 1–2 teeth or lobes.

all basal, new leaves tightly clustered;

blades spatulate or oblanceolate to linear-oblanceolate, entire, glabrous or ± hairy, often lanuginose, sericeous, or strigoso-canescent, eglandular or ± gland-dotted.

Peduncles

8–29 cm, ± hairy.

0.5–30 cm, ± hairy.

Involucres

5–10 × 7–15 mm.

7–12 × 8–16 mm.

Ray florets

9–25;

corollas 8.2–16.8 mm.

8–15(–21);

corollas 9–19 mm.

Disc florets

50–200+;

corollas yellow, 1.6–3 mm.

25–200+;

corollas yellow, 2.7–4.3 mm.

Outer phyllaries

8–21, 2.4–5.5 mm, margins 0–0.2 mm wide, not scarious or scarious, abaxial faces ± hairy.

6–12, 3.9–9(–11.5) mm, margins 0–0.4 mm wide, sometimes slightly scarious, abaxial faces ± hairy.

Heads

8–50(–80) per plant, borne singly or in corymbiform arrays.

1–35(–60) per plant, borne singly.

Cypselae

1.5–2.6 mm;

pappi of 4–8 obovate, often aristate scales 1–2.5 mm.

2–4 mm;

pappi of 5–8 obovate to oblanceolate, aristate scales 2.2–3.7 mm.

Tetraneuris linearifolia

Tetraneuris acaulis

Distribution
from FNA
KS; NM; OK; TX; n Mexico
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[BONAP county map]
from FNA
AZ; CA; CO; ID; KS; MT; ND; NE; NM; NV; OK; SD; TX; UT; WY; AB; SK
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[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

Plants of Tetraneuris linearifolia morphologically intermediate between the two varieties are found in Bee, Duval, Live Oak, and McMullen counties, Texas.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 4 (4 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Stems erect, branched distally; leaves ± hairy, basal blades entire or with 2(–6) teeth or lobes, proximal cauline blades entire or with 1–2 teeth or lobes, mid blades (0.9–)1.4–3(–4.8) mm wide; involucres (7–)8–11(–12) mm diam.; pappus scales aristate; Kansas, New Mexico, Oklahoma, Texas, on limestone-derived soils
var. linearifolia
1. Stems erect or ± decumbent, ± branched distally; leaves usually densely hairy, basal blades with 2–6 teeth or lobes, proximal cauline blades with 2(–6) teeth or lobes, mid blades (2.5–)3.9–9.5(–16) mm wide; involucres 12–15 mm diam.; pappus scales not aristate; s Texas on sand
var. arenicola
1. Leaves sparsely to densely hairy, often lanuginose, sometimes sericeous, not strigoso-canescent; peduncles 0.5–8(–12) cm; outer phyllaries 6.8–9(–11.5) mm; growing at (2100–)3000–3900 m
var. caespitosa
1. Leaves sparsely to densely hairy, often strigoso-canescent; peduncles (1–)5–20(–30) cm; outer phyllaries 4–7.5 mm; growing at 700–2900(–3500) m
→ 2
2. Leaf blades usually densely, sometimes sparsely, strigoso-canescent; e of continental divide (except for 2 reports from nw Idaho)
var. acaulis
2. Leaf blades glabrous or sparsely to moderately to sometimes densely hairy, not strigoso-canescent; w of continental divide
→ 3
3. Leaf blades glabrous or usually sparsely to moderately or sometimes densely hairy, densely gland-dotted; n Arizona, se California, w Colorado, s Idaho, e Nevada, Utah
var. arizonica
3. Leaf blades glabrous or sparsely to moderately hairy, at least some (usually most or all) eglandular or sparsely gland-dotted; w Colorado, c, ne Utah, and sw Wyoming
var. epunctata
Source FNA vol. 21, p. 448. FNA vol. 21, p. 451.
Parent taxa Asteraceae > tribe Heliantheae > subtribe Gaillardiinae > Tetraneuris Asteraceae > tribe Heliantheae > subtribe Gaillardiinae > Tetraneuris
Sibling taxa
T. acaulis, T. argentea, T. herbacea, T. ivesiana, T. scaposa, T. torreyana, T. turneri, T. verdiensis
T. argentea, T. herbacea, T. ivesiana, T. linearifolia, T. scaposa, T. torreyana, T. turneri, T. verdiensis
Subordinate taxa
T. linearifolia var. arenicola, T. linearifolia var. linearifolia
T. acaulis var. acaulis, T. acaulis var. arizonica, T. acaulis var. caespitosa, T. acaulis var. epunctata
Synonyms Hymenoxys linearifolia Gaillardia acaulis, Hymenoxys acaulis
Name authority (Hooker) Greene: Pittonia 3: 269. (1898) (Pursh) Greene: Pittonia 3: 265. (1898)
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