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Ives' four-nerve-daisy

fineleaf fournerve daisy, slimleaf actinea

Habit Perennials, 10–26+ cm. Annuals, 16–50+ cm.
Caudices

± branched, branches notably thickened distally.

none.

Stems

(1–)5–20(–30), erect, leafy, ± branched distally.

1–10, erect or ± decumbent, leafy, unbranched or branched distally.

Leaves

basal and cauline, entire, glabrous or sparsely to moderately lanuginose to sericeous, usually densely gland-dotted;

basal leaves tightly clustered, blades linear-oblanceolate;

proximal cauline blades linear-oblanceolate;

mid blades linear-oblanceolate to linear, 1.7–2 mm wide;

distal blades linear-oblanceolate to linear.

basal and cauline, ± hairy, ± gland-dotted;

basal blades spatulate to oblanceolate, entire or with 2–6 teeth or lobes;

proximal cauline blades narrowly spatulate or oblanceolate to linear-oblanceolate, entire or with 2–6 teeth or lobes;

mid blades oblanceolate to linear-oblanceolate to linear, 0.9–9.5(–16) mm wide, entire or with 1–2 teeth or lobes;

distal blades narrowly oblanceolate to linear-oblanceolate to linear, entire or with 1–2 teeth or lobes.

Peduncles

5–18.5 cm, ± hairy, usually densely hairy distally.

8–29 cm, ± hairy.

Involucres

8–12 × 13–17 mm.

5–10 × 7–15 mm.

Ray florets

7–10;

corollas 10–20 mm.

9–25;

corollas 8.2–16.8 mm.

Disc florets

(40–)80–150+;

corollas yellow, 3.3–4.5 mm.

50–200+;

corollas yellow, 1.6–3 mm.

Outer phyllaries

7–12, 5–6.4 mm, margins 0–0.3(–0.4) mm wide, usually scarious, abaxial faces ± hairy.

8–21, 2.4–5.5 mm, margins 0–0.2 mm wide, not scarious or scarious, abaxial faces ± hairy.

Heads

(1–)5–30(–40) per plant (1–5 per stem), borne singly or in paniculiform to corymbiform arrays.

8–50(–80) per plant, borne singly or in corymbiform arrays.

Cypselae

3–4.1 mm;

pappi of 5–7 lanceolate or obovate to oblanceolate, aristate scales 2.7–4.5 mm.

1.5–2.6 mm;

pappi of 4–8 obovate, often aristate scales 1–2.5 mm.

2n

= 28, 30, 56, 60.

Tetraneuris ivesiana

Tetraneuris linearifolia

Phenology Flowering Apr–Aug.
Habitat Roadsides, hillsides, open treeless areas, edges of woods
Elevation 1300–2900 m (4300–9500 ft)
Distribution
from FNA
AZ; CO; NM; UT
[WildflowerSearch map]
[BONAP county map]
from FNA
KS; NM; OK; TX; n Mexico
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

Plants of Tetraneuris linearifolia morphologically intermediate between the two varieties are found in Bee, Duval, Live Oak, and McMullen counties, Texas.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Stems erect, branched distally; leaves ± hairy, basal blades entire or with 2(–6) teeth or lobes, proximal cauline blades entire or with 1–2 teeth or lobes, mid blades (0.9–)1.4–3(–4.8) mm wide; involucres (7–)8–11(–12) mm diam.; pappus scales aristate; Kansas, New Mexico, Oklahoma, Texas, on limestone-derived soils
var. linearifolia
1. Stems erect or ± decumbent, ± branched distally; leaves usually densely hairy, basal blades with 2–6 teeth or lobes, proximal cauline blades with 2(–6) teeth or lobes, mid blades (2.5–)3.9–9.5(–16) mm wide; involucres 12–15 mm diam.; pappus scales not aristate; s Texas on sand
var. arenicola
Source FNA vol. 21, p. 450. FNA vol. 21, p. 448.
Parent taxa Asteraceae > tribe Heliantheae > subtribe Gaillardiinae > Tetraneuris Asteraceae > tribe Heliantheae > subtribe Gaillardiinae > Tetraneuris
Sibling taxa
T. acaulis, T. argentea, T. herbacea, T. linearifolia, T. scaposa, T. torreyana, T. turneri, T. verdiensis
T. acaulis, T. argentea, T. herbacea, T. ivesiana, T. scaposa, T. torreyana, T. turneri, T. verdiensis
Subordinate taxa
T. linearifolia var. arenicola, T. linearifolia var. linearifolia
Synonyms Hymenoxys acaulis var. ivesiana, Hymenoxys argentea var. ivesiana, Hymenoxys ivesiana, T. intermedia, T. mancosensis, T. pilosa Hymenoxys linearifolia
Name authority Greene: Pittonia 3: 269. (1898) (Hooker) Greene: Pittonia 3: 269. (1898)
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