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Ives' four-nerve-daisy

perky sue, stemless four-nerve daisy, stemless hymenoxys, stemless woollybase

Habit Perennials, 10–26+ cm. Perennials, 2–30+ cm.
Caudices

± branched, branches notably thickened distally.

± branched, branches notably thickened distally.

Stems

(1–)5–20(–30), erect, leafy, ± branched distally.

1–35(–60), erect, unbranched.

Leaves

basal and cauline, entire, glabrous or sparsely to moderately lanuginose to sericeous, usually densely gland-dotted;

basal leaves tightly clustered, blades linear-oblanceolate;

proximal cauline blades linear-oblanceolate;

mid blades linear-oblanceolate to linear, 1.7–2 mm wide;

distal blades linear-oblanceolate to linear.

all basal, new leaves tightly clustered;

blades spatulate or oblanceolate to linear-oblanceolate, entire, glabrous or ± hairy, often lanuginose, sericeous, or strigoso-canescent, eglandular or ± gland-dotted.

Peduncles

5–18.5 cm, ± hairy, usually densely hairy distally.

0.5–30 cm, ± hairy.

Involucres

8–12 × 13–17 mm.

7–12 × 8–16 mm.

Ray florets

7–10;

corollas 10–20 mm.

8–15(–21);

corollas 9–19 mm.

Disc florets

(40–)80–150+;

corollas yellow, 3.3–4.5 mm.

25–200+;

corollas yellow, 2.7–4.3 mm.

Outer phyllaries

7–12, 5–6.4 mm, margins 0–0.3(–0.4) mm wide, usually scarious, abaxial faces ± hairy.

6–12, 3.9–9(–11.5) mm, margins 0–0.4 mm wide, sometimes slightly scarious, abaxial faces ± hairy.

Heads

(1–)5–30(–40) per plant (1–5 per stem), borne singly or in paniculiform to corymbiform arrays.

1–35(–60) per plant, borne singly.

Cypselae

3–4.1 mm;

pappi of 5–7 lanceolate or obovate to oblanceolate, aristate scales 2.7–4.5 mm.

2–4 mm;

pappi of 5–8 obovate to oblanceolate, aristate scales 2.2–3.7 mm.

2n

= 28, 30, 56, 60.

Tetraneuris ivesiana

Tetraneuris acaulis

Phenology Flowering Apr–Aug.
Habitat Roadsides, hillsides, open treeless areas, edges of woods
Elevation 1300–2900 m (4300–9500 ft)
Distribution
from FNA
AZ; CO; NM; UT
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; CO; ID; KS; MT; ND; NE; NM; NV; OK; SD; TX; UT; WY; AB; SK
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[BONAP county map]
Discussion

Varieties 4 (4 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaves sparsely to densely hairy, often lanuginose, sometimes sericeous, not strigoso-canescent; peduncles 0.5–8(–12) cm; outer phyllaries 6.8–9(–11.5) mm; growing at (2100–)3000–3900 m
var. caespitosa
1. Leaves sparsely to densely hairy, often strigoso-canescent; peduncles (1–)5–20(–30) cm; outer phyllaries 4–7.5 mm; growing at 700–2900(–3500) m
→ 2
2. Leaf blades usually densely, sometimes sparsely, strigoso-canescent; e of continental divide (except for 2 reports from nw Idaho)
var. acaulis
2. Leaf blades glabrous or sparsely to moderately to sometimes densely hairy, not strigoso-canescent; w of continental divide
→ 3
3. Leaf blades glabrous or usually sparsely to moderately or sometimes densely hairy, densely gland-dotted; n Arizona, se California, w Colorado, s Idaho, e Nevada, Utah
var. arizonica
3. Leaf blades glabrous or sparsely to moderately hairy, at least some (usually most or all) eglandular or sparsely gland-dotted; w Colorado, c, ne Utah, and sw Wyoming
var. epunctata
Source FNA vol. 21, p. 450. FNA vol. 21, p. 451.
Parent taxa Asteraceae > tribe Heliantheae > subtribe Gaillardiinae > Tetraneuris Asteraceae > tribe Heliantheae > subtribe Gaillardiinae > Tetraneuris
Sibling taxa
T. acaulis, T. argentea, T. herbacea, T. linearifolia, T. scaposa, T. torreyana, T. turneri, T. verdiensis
T. argentea, T. herbacea, T. ivesiana, T. linearifolia, T. scaposa, T. torreyana, T. turneri, T. verdiensis
Subordinate taxa
T. acaulis var. acaulis, T. acaulis var. arizonica, T. acaulis var. caespitosa, T. acaulis var. epunctata
Synonyms Hymenoxys acaulis var. ivesiana, Hymenoxys argentea var. ivesiana, Hymenoxys ivesiana, T. intermedia, T. mancosensis, T. pilosa Gaillardia acaulis, Hymenoxys acaulis
Name authority Greene: Pittonia 3: 269. (1898) (Pursh) Greene: Pittonia 3: 265. (1898)
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