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perky sue, stemless four-nerve daisy, stemless hymenoxys, stemless woollybase

fineleaf fournerve daisy, slimleaf actinea

Habit Perennials, 2–30+ cm. Annuals, 16–50+ cm.
Caudices

± branched, branches notably thickened distally.

none.

Stems

1–35(–60), erect, unbranched.

1–10, erect or ± decumbent, leafy, unbranched or branched distally.

Leaves

all basal, new leaves tightly clustered;

blades spatulate or oblanceolate to linear-oblanceolate, entire, glabrous or ± hairy, often lanuginose, sericeous, or strigoso-canescent, eglandular or ± gland-dotted.

basal and cauline, ± hairy, ± gland-dotted;

basal blades spatulate to oblanceolate, entire or with 2–6 teeth or lobes;

proximal cauline blades narrowly spatulate or oblanceolate to linear-oblanceolate, entire or with 2–6 teeth or lobes;

mid blades oblanceolate to linear-oblanceolate to linear, 0.9–9.5(–16) mm wide, entire or with 1–2 teeth or lobes;

distal blades narrowly oblanceolate to linear-oblanceolate to linear, entire or with 1–2 teeth or lobes.

Peduncles

0.5–30 cm, ± hairy.

8–29 cm, ± hairy.

Involucres

7–12 × 8–16 mm.

5–10 × 7–15 mm.

Ray florets

8–15(–21);

corollas 9–19 mm.

9–25;

corollas 8.2–16.8 mm.

Disc florets

25–200+;

corollas yellow, 2.7–4.3 mm.

50–200+;

corollas yellow, 1.6–3 mm.

Outer phyllaries

6–12, 3.9–9(–11.5) mm, margins 0–0.4 mm wide, sometimes slightly scarious, abaxial faces ± hairy.

8–21, 2.4–5.5 mm, margins 0–0.2 mm wide, not scarious or scarious, abaxial faces ± hairy.

Heads

1–35(–60) per plant, borne singly.

8–50(–80) per plant, borne singly or in corymbiform arrays.

Cypselae

2–4 mm;

pappi of 5–8 obovate to oblanceolate, aristate scales 2.2–3.7 mm.

1.5–2.6 mm;

pappi of 4–8 obovate, often aristate scales 1–2.5 mm.

Tetraneuris acaulis

Tetraneuris linearifolia

Distribution
from FNA
AZ; CA; CO; ID; KS; MT; ND; NE; NM; NV; OK; SD; TX; UT; WY; AB; SK
[WildflowerSearch map]
[BONAP county map]
from FNA
KS; NM; OK; TX; n Mexico
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 4 (4 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

Plants of Tetraneuris linearifolia morphologically intermediate between the two varieties are found in Bee, Duval, Live Oak, and McMullen counties, Texas.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaves sparsely to densely hairy, often lanuginose, sometimes sericeous, not strigoso-canescent; peduncles 0.5–8(–12) cm; outer phyllaries 6.8–9(–11.5) mm; growing at (2100–)3000–3900 m
var. caespitosa
1. Leaves sparsely to densely hairy, often strigoso-canescent; peduncles (1–)5–20(–30) cm; outer phyllaries 4–7.5 mm; growing at 700–2900(–3500) m
→ 2
2. Leaf blades usually densely, sometimes sparsely, strigoso-canescent; e of continental divide (except for 2 reports from nw Idaho)
var. acaulis
2. Leaf blades glabrous or sparsely to moderately to sometimes densely hairy, not strigoso-canescent; w of continental divide
→ 3
3. Leaf blades glabrous or usually sparsely to moderately or sometimes densely hairy, densely gland-dotted; n Arizona, se California, w Colorado, s Idaho, e Nevada, Utah
var. arizonica
3. Leaf blades glabrous or sparsely to moderately hairy, at least some (usually most or all) eglandular or sparsely gland-dotted; w Colorado, c, ne Utah, and sw Wyoming
var. epunctata
1. Stems erect, branched distally; leaves ± hairy, basal blades entire or with 2(–6) teeth or lobes, proximal cauline blades entire or with 1–2 teeth or lobes, mid blades (0.9–)1.4–3(–4.8) mm wide; involucres (7–)8–11(–12) mm diam.; pappus scales aristate; Kansas, New Mexico, Oklahoma, Texas, on limestone-derived soils
var. linearifolia
1. Stems erect or ± decumbent, ± branched distally; leaves usually densely hairy, basal blades with 2–6 teeth or lobes, proximal cauline blades with 2(–6) teeth or lobes, mid blades (2.5–)3.9–9.5(–16) mm wide; involucres 12–15 mm diam.; pappus scales not aristate; s Texas on sand
var. arenicola
Source FNA vol. 21, p. 451. FNA vol. 21, p. 448.
Parent taxa Asteraceae > tribe Heliantheae > subtribe Gaillardiinae > Tetraneuris Asteraceae > tribe Heliantheae > subtribe Gaillardiinae > Tetraneuris
Sibling taxa
T. argentea, T. herbacea, T. ivesiana, T. linearifolia, T. scaposa, T. torreyana, T. turneri, T. verdiensis
T. acaulis, T. argentea, T. herbacea, T. ivesiana, T. scaposa, T. torreyana, T. turneri, T. verdiensis
Subordinate taxa
T. acaulis var. acaulis, T. acaulis var. arizonica, T. acaulis var. caespitosa, T. acaulis var. epunctata
T. linearifolia var. arenicola, T. linearifolia var. linearifolia
Synonyms Gaillardia acaulis, Hymenoxys acaulis Hymenoxys linearifolia
Name authority (Pursh) Greene: Pittonia 3: 265. (1898) (Hooker) Greene: Pittonia 3: 269. (1898)
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