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perky sue, stemless four-nerve daisy, stemless hymenoxys, stemless woollybase

Photo is of parent taxon

caespitose four-nerve daisy

Habit Perennials, 2–30+ cm. Plants 2–8(–12+) cm.
Caudices

± branched, branches notably thickened distally.

Stems

1–35(–60), erect, unbranched.

Leaves

all basal, new leaves tightly clustered;

blades spatulate or oblanceolate to linear-oblanceolate, entire, glabrous or ± hairy, often lanuginose, sericeous, or strigoso-canescent, eglandular or ± gland-dotted.

blades spatulate or oblanceolate to linear-oblanceolate, sparsely to densely hairy, often lanuginose, sometimes sericeous, sparsely to sometimes moderately or densely gland-dotted.

Peduncles

0.5–30 cm, ± hairy.

0.5–8(–12) cm.

Involucres

7–12 × 8–16 mm.

Ray florets

8–15(–21);

corollas 9–19 mm.

9–15(–21);

corollas 9–13 mm.

Disc florets

25–200+;

corollas yellow, 2.7–4.3 mm.

Outer phyllaries

6–12, 3.9–9(–11.5) mm, margins 0–0.4 mm wide, sometimes slightly scarious, abaxial faces ± hairy.

6–12, 6.8–9(–11.5) mm, margins 0–0.2 mm wide, sometimes slightly scarious, abaxial faces usually densely hairy.

Heads

1–35(–60) per plant, borne singly.

1–35(–60) per plant.

Cypselae

2–4 mm;

pappi of 5–8 obovate to oblanceolate, aristate scales 2.2–3.7 mm.

2–3(–4) mm;

pappi 2.5–3.5 mm.

2n

= 28, 30.

Tetraneuris acaulis

Tetraneuris acaulis var. caespitosa

Phenology Flowering (May–)Jun–Aug(–Sep).
Habitat Edges of montane forests, alpine meadows above timberline, open savannas
Elevation (2100–)3000–3900 m ((6900–)9800–12800 ft)
Distribution
from FNA
AZ; CA; CO; ID; KS; MT; ND; NE; NM; NV; OK; SD; TX; UT; WY; AB; SK
[WildflowerSearch map]
[BONAP county map]
from FNA
CO; NM; WY
[BONAP county map]
Discussion

Varieties 4 (4 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaves sparsely to densely hairy, often lanuginose, sometimes sericeous, not strigoso-canescent; peduncles 0.5–8(–12) cm; outer phyllaries 6.8–9(–11.5) mm; growing at (2100–)3000–3900 m
var. caespitosa
1. Leaves sparsely to densely hairy, often strigoso-canescent; peduncles (1–)5–20(–30) cm; outer phyllaries 4–7.5 mm; growing at 700–2900(–3500) m
→ 2
2. Leaf blades usually densely, sometimes sparsely, strigoso-canescent; e of continental divide (except for 2 reports from nw Idaho)
var. acaulis
2. Leaf blades glabrous or sparsely to moderately to sometimes densely hairy, not strigoso-canescent; w of continental divide
→ 3
3. Leaf blades glabrous or usually sparsely to moderately or sometimes densely hairy, densely gland-dotted; n Arizona, se California, w Colorado, s Idaho, e Nevada, Utah
var. arizonica
3. Leaf blades glabrous or sparsely to moderately hairy, at least some (usually most or all) eglandular or sparsely gland-dotted; w Colorado, c, ne Utah, and sw Wyoming
var. epunctata
Source FNA vol. 21, p. 451. FNA vol. 21, p. 452.
Parent taxa Asteraceae > tribe Heliantheae > subtribe Gaillardiinae > Tetraneuris Asteraceae > tribe Heliantheae > subtribe Gaillardiinae > Tetraneuris > Tetraneuris acaulis
Sibling taxa
T. argentea, T. herbacea, T. ivesiana, T. linearifolia, T. scaposa, T. torreyana, T. turneri, T. verdiensis
T. acaulis var. acaulis, T. acaulis var. arizonica, T. acaulis var. epunctata
Subordinate taxa
T. acaulis var. acaulis, T. acaulis var. arizonica, T. acaulis var. caespitosa, T. acaulis var. epunctata
Synonyms Gaillardia acaulis, Hymenoxys acaulis Hymenoxys acaulis var. caespitosa, T. brevifolia, T. lanata, T. lanigera
Name authority (Pursh) Greene: Pittonia 3: 265. (1898) A. Nelson: Bot. Gaz. 28: 127. (1899)
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