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perky sue, stemless four-nerve daisy, stemless hymenoxys, stemless woollybase

Photo is of parent taxon

stemless four-nerve daisy

Habit Perennials, 2–30+ cm. Plants (3–)10–20(–30+) cm.
Caudices

± branched, branches notably thickened distally.

Stems

1–35(–60), erect, unbranched.

Leaves

all basal, new leaves tightly clustered;

blades spatulate or oblanceolate to linear-oblanceolate, entire, glabrous or ± hairy, often lanuginose, sericeous, or strigoso-canescent, eglandular or ± gland-dotted.

blades spatulate or oblanceolate to linear-oblanceolate, usually densely, sometimes sparsely, strigoso-canescent, sparsely to moderately or densely gland-dotted.

Peduncles

0.5–30 cm, ± hairy.

(2–)8–20(–30) cm.

Involucres

7–12 × 8–16 mm.

Ray florets

8–15(–21);

corollas 9–19 mm.

8–14;

corollas 9–15 mm.

Disc florets

25–200+;

corollas yellow, 2.7–4.3 mm.

Outer phyllaries

6–12, 3.9–9(–11.5) mm, margins 0–0.4 mm wide, sometimes slightly scarious, abaxial faces ± hairy.

6–10, 4–7 mm, margins 0–0.2(–0.4) mm wide, sometimes slightly scarious, abaxial faces usually densely hairy.

Heads

1–35(–60) per plant, borne singly.

1–10(–15) per plant.

Cypselae

2–4 mm;

pappi of 5–8 obovate to oblanceolate, aristate scales 2.2–3.7 mm.

2.5–3 mm;

pappi 2.2–2.9 mm.

2n

= 28, 30, 56, 60.

Tetraneuris acaulis

Tetraneuris acaulis var. acaulis

Phenology Flowering (Apr–)May–Jul(–Oct).
Habitat Roadsides, hillsides, grasslands, edges of woods
Elevation 700–2000(–3500) m (2300–6600(–11500) ft)
Distribution
from FNA
AZ; CA; CO; ID; KS; MT; ND; NE; NM; NV; OK; SD; TX; UT; WY; AB; SK
[WildflowerSearch map]
[BONAP county map]
from FNA
CO; ID; KS; MT; ND; NE; NM; OK; SD; TX; WY; AB; SK
[BONAP county map]
Discussion

Varieties 4 (4 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Some plants of var. acaulis from Colorado and southeastern Wyoming have considerably less hairy and/or more densely gland-dotted leaves, making them easy to confuse with var. arizonica. As to distribution, label on a collection at LL reads, “From W. A. Curtis, Lewistown [Idaho], without definite locality but probably collected in that vicinity.” Also, the holotype and isotype of Tetraneuris septentrionalis were collected in “Palouse County, Idaho.” These are the only collections we have seen that place var. acaulis west of the continental divide.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaves sparsely to densely hairy, often lanuginose, sometimes sericeous, not strigoso-canescent; peduncles 0.5–8(–12) cm; outer phyllaries 6.8–9(–11.5) mm; growing at (2100–)3000–3900 m
var. caespitosa
1. Leaves sparsely to densely hairy, often strigoso-canescent; peduncles (1–)5–20(–30) cm; outer phyllaries 4–7.5 mm; growing at 700–2900(–3500) m
→ 2
2. Leaf blades usually densely, sometimes sparsely, strigoso-canescent; e of continental divide (except for 2 reports from nw Idaho)
var. acaulis
2. Leaf blades glabrous or sparsely to moderately to sometimes densely hairy, not strigoso-canescent; w of continental divide
→ 3
3. Leaf blades glabrous or usually sparsely to moderately or sometimes densely hairy, densely gland-dotted; n Arizona, se California, w Colorado, s Idaho, e Nevada, Utah
var. arizonica
3. Leaf blades glabrous or sparsely to moderately hairy, at least some (usually most or all) eglandular or sparsely gland-dotted; w Colorado, c, ne Utah, and sw Wyoming
var. epunctata
Source FNA vol. 21, p. 451. FNA vol. 21, p. 452.
Parent taxa Asteraceae > tribe Heliantheae > subtribe Gaillardiinae > Tetraneuris Asteraceae > tribe Heliantheae > subtribe Gaillardiinae > Tetraneuris > Tetraneuris acaulis
Sibling taxa
T. argentea, T. herbacea, T. ivesiana, T. linearifolia, T. scaposa, T. torreyana, T. turneri, T. verdiensis
T. acaulis var. arizonica, T. acaulis var. caespitosa, T. acaulis var. epunctata
Subordinate taxa
T. acaulis var. acaulis, T. acaulis var. arizonica, T. acaulis var. caespitosa, T. acaulis var. epunctata
Synonyms Gaillardia acaulis, Hymenoxys acaulis T. eradiata, T. incana, T. pygmaea, T. septentrionalis, T. simplex
Name authority (Pursh) Greene: Pittonia 3: 265. (1898) unknown
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