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pineland nerveray

nerveray

Habit Perennials, mostly 30–120 cm.
Stems

patently to retrorsely ± villous.

erect.

Leaves

cauline, not connate-perfoliate;

blades ovate to rhombic or lanceolate, 7–20 × 3–13 cm, coarsely toothed.

basal and/or cauline; mostly opposite; petiolate, subpetiolate, or sessile (bases sometimes connate-perfoliate);

blades lanceolate, ovate, rhombic, or rounded-deltate, sometimes pinnatifid, ultimate margins usually toothed, faces glabrate, sparsely hispidulous, or puberulent, gland-dotted.

Involucres

obpyramidal to hemispheric, 12–25+ mm diam.

Receptacles

conic, paleate (paleae persistent, lanceolate to lance-ovate, flat or weakly conduplicate, apices acute).

Ray florets

6–14;

laminae 15–40+ mm.

6–21+, pistillate, fertile;

corollas yellow (often with reddish nerves).

Disc florets

25–150+, bisexual, fertile;

corollas yellow, tubes (basally dilated) much shorter than ampliate, cylindric throats, lobes 5, deltate.

Phyllaries

persistent, 10–25+ in ± 2 series (outer 4 broadly lanceolate, foliaceous, the inner ovate to lanceolate, smaller, more scarious, each subtending a ray floret).

Heads

radiate, borne singly or in loose, corymbiform arrays.

Cypselae

± ovoid, 4–6 mm;

pappi 0.

± ovoid or plumply 4- or 5-angled, finely 32–40-ribbed, sparsely strigose or glabrous;

pappi 0, or of 1–10+ subulate to acerose scales (to 0.5 mm), or of 16–30, ± spatulate scales (0.5–2 mm).

x

= 17.

2n

= 34.

Tetragonotheca helianthoides

Tetragonotheca

Phenology Flowering mid spring–early summer.
Habitat Sandy soils, often in thickets
Elevation 0–400 m (0–1300 ft)
Distribution
from FNA
AL; FL; GA; MS; NC; SC; TN; VA
[WildflowerSearch map]
[BONAP county map]
from USDA
n Mexico; s United States
[BONAP county map]
Discussion

Species 4 (4 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Pappi of 16–30, ± obovate to spatulate scales 0.5–2 mm
→ 2
1. Pappi 0, or of 1–10+ subulate to acerose scales to 0.5 mm
→ 3
2. Leaves mostly cauline (distal connate-perfoliate); ray florets 12–13, laminae 8–18 mm
T. ludoviciana
2. Leaves mostly basal (cauline much smaller, none connate-perfoliate); ray florets 16–21+, laminae 20–30+ mm
T. repanda
3. Stems sparsely tomentulose or glabrous (nodes usually auriculate-appendaged); ray laminae 10–18 mm; cypselae ± quadrate, 2.5–4 mm
T. texana
3. Stems patently to retrorsely ± villous; ray laminae 20–40+ mm; cypselae ± ovoid, 4–6 mm
T. helianthoides
Source FNA vol. 21, p. 179. FNA vol. 21, p. 178. Author: John L. Strother.
Parent taxa Asteraceae > tribe Heliantheae > subtribe Galinsoginae > Tetragonotheca Asteraceae > tribe Heliantheae > subtribe Galinsoginae
Sibling taxa
T. ludoviciana, T. repanda, T. texana
Subordinate taxa
T. helianthoides, T. ludoviciana, T. repanda, T. texana
Name authority Linnaeus: Sp. Pl. 2: 903. (1753) Linnaeus: Sp. Pl. 2: 903. (1753): Gen. Pl. ed. 5, 384. (1754)
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