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Nuttall's horsebrush

cottonthorn, horse-brush

Habit Shrubs, 10–120 cm. Shrubs, mostly 30–200 cm.
Stems

1–5+, erect, spiny, pannose but for glabrescent streaks.

1–5+, erect to spreading (tomentose to pannose or glabrous, often spiny).

Leaves

primaries forming straight or recurved spines, 5–25 mm;

secondaries spatulate, 10–20 mm, tomentose to nearly glabrous.

(usually with fascicles of secondary leaves in axils of the primary; primaries sometimes becoming spines) cauline; alternate;

sessile (or obscurely petiolate);

blades obscurely palmately nerved (usually 1-nerved), linear to filiform, narrowly lanceolate, or oblanceolate, margins entire, faces glabrous or lanate to tomentose, sometimes glabrescent.

Peduncles

2–12 mm.

Involucres

turbinate to cylindric, 6–9 mm.

turbinate to cylindric or hemispheric, 3–10+ mm diam.

Receptacles

flat, foveolate, epaleate.

Florets

4;

corollas bright yellow, 8–10 mm.

Ray florets

0.

Disc florets

4–9, bisexual, fertile;

corollas cream to bright yellow, tubes longer than funnelform or abruptly dilated throats, lobes 5, recurved, linear to lanceolate;

style branches stigmatic in 2 lines, apices usually truncate to rounded-truncate, sometimes truncate-penicillate (appendages essentially 0).

Phyllaries

4, oblong.

persistent, 4–6 in 1–2 series, erect, distinct, mostly oblong to ovate or lanceolate, equal or subequal, margins obscurely scarious.

Calyculi

0.

Heads

4–6.

discoid, (1–3) in distil axils or (3–8) in corymbiform clusters.

Cypselae

4–6 mm, densely hirsute;

pappi of 75–100 bristles 9–10 mm.

prismatic to obconic or fusiform, obscurely 5-ribbed, glabrous or hirsute to densely pilose (hairs often obscuring pappi);

pappi 0 or of 70–150 bristles or of 20–30, white to stramineous, subulate to setiform scales.

x

= 30.

2n

= 60.

Tetradymia nuttallii

Tetradymia

Phenology Flowering spring–summer.
Habitat Rocky or sandy places, sagebrush scrub or shadscale scrub
Elevation 1300–2100 m (4300–6900 ft)
Distribution
from FNA
NV; UT; WY
[WildflowerSearch map]
[BONAP county map]
from USDA
w North America; nw Mexico
[BONAP county map]
Discussion

Species 10 (10 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Indumentum of stems evenly pannose; cypselae with white hairs that obscure or replace pappi; pappi 0 or of 20–30, subulate to setiform scales
→ 2
1. Indumentum of stems interrupted by linear, tomentose to nearly glabrous streaks; cypselae glabrous or with induments shorter than cypselae, not obscuring or replacing pappi; pappi of 70–150 bristles
→ 5
2. Primary leaves forming rigid spines; heads in clusters of 1–3 (in axils of spines)
→ 3
2. Primary leaves flaccid or stiff and pungent, not forming spines; heads in corymbiform clusters of 3–7
→ 4
3. Spines straight, 10–50 mm, glabrescent; involucres mostly turbinate, 7–9 mm; cypselae 4–5 mm
T. axillaris
3. Spines recurved, 5–25 mm, tomentose; involucres mostly campanulate, 8–12 mm; cypselae 6–8 mm
T. spinosa
4. Blades of primary leaves lance-linear to spatulate, becoming stiff, pungent; florets 5–8; cypselae 6–8 mm, epappose
T. comosa
4. Blades of primary leaves filiform, flaccid; florets 4(–5); cypselae 4–6 mm, pappose
T. tetrameres
5. Primary leaves not forming spines
→ 6
5. Primary leaves becoming rigid, ± persistent spines
→ 8
6. Blades of primary leaves subulate and ± appressed to stems, contrasting with subtended clavate, glabrous or glabrescent blades of secondary leaves
T. glabrata
6. Blades of primary leaves filiform to spatulate, spreading, similar to subtended tomentose to sericeous blades of secondary leaves
→ 7
7. Blades of primary leaves spatulate to lanceolate; pappus bristles weakly, if at all, distally dilated
T. canescens
7. Blades of primary leaves linear to filiform; pappus bristles distally dilated
T. filifolia
8. Blades of secondary leaves filiform to clavate; cypselae glabrous
T. argyraea
8. Blades of secondary leaves spatulate; cypselae densely hirsute or villous to densely pilose
→ 9
9. Blades of secondary leaves greenish, usually villous, sometimes glabrescent; florets mostly 4; cypselae 4–6 mm, densely hirsute
T. nuttallii
9. Blades of secondary leaves silvery sericeous; florets mostly 5; cypselae 6–8 mm, villous to densely pilose
T. stenolepis
Source FNA vol. 20, p. 631. FNA vol. 20, p. 629. Author: John L. Strother.
Parent taxa Asteraceae > tribe Senecioneae > Tetradymia Asteraceae > tribe Senecioneae
Sibling taxa
T. argyraea, T. axillaris, T. canescens, T. comosa, T. filifolia, T. glabrata, T. spinosa, T. stenolepis, T. tetrameres
Subordinate taxa
T. argyraea, T. axillaris, T. canescens, T. comosa, T. filifolia, T. glabrata, T. nuttallii, T. spinosa, T. stenolepis, T. tetrameres
Name authority Torrey & A. Gray: Fl. N. Amer. 2: 447. (1843) de Candolle: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 6: 440. (1838)
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