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halberd fern

broad halberd fern

Habit Plants terrestrial or on rock.
Stems

short-creeping to erect, stolons absent.

erect, compact.

Leaves

monomorphic, evergreen.

Petiole

3/4–3 times length of blade, base not swollen;

vascular bundles more than 3, arranged in an arc, ± round in cross section.

straw-colored or rarely brown, 1–2 times length of blade, glabrous on both surfaces, base scaly;

scales linear to lanceolate, 3–10 × 0.5–2 mm.

Blade

lanceolate to deltate or pentagonal, entire to 1-pinnate-pinnatifid [3-pinnate-pinnatifid], reduced distally to shallowly lobed or hastate apex, herbaceous to papery.

ovate to pentagonal, 1-pinnate or (in small plants) ternately lobed, (12–)20–40(–50) × 14–40(–45) cm.

Pinnae

not articulate to rachis, segment margins entire to sinuate or shallowly lobed;

proximal pinnae not or only slightly reduced, sessile to short-petiolulate, base equilateral or often inequilateral with prominent basiscopic lobe(s);

costae adaxially rounded or shallowly grooved, grooves not continuous from rachis to costae;

indument lacking or of multicellular hairs on costae abaxially, of multicellular hairs on costae adaxially.

1–2(–3) pairs, 12–22 cm, base cordate, margins shallowly lobed, with pair of basal lobes and without proliferous buds in axils.

Veins

reticulate, areoles with or without included veinlets.

Indusia

peltate.

Sori

in 1–several rows between midrib and margin, round;

indusia peltate to round-reniform and with narrow sinus, persistent or caducous.

Spores

brownish, with inflated folds or wings.

Rachis

and costae glabrous abaxially.

Areoles

with included veinlets.

x

= 40.

2n

= 160.

Tectaria

Tectaria heracleifolia

Habitat Rocky hammocks, limestone outcrops in shade, cave entrances
Elevation 0–100 m (0–300 ft)
Distribution
from USDA
Mostly tropical
[BONAP county map]
from FNA
FL; TX; Mexico; Central America; West Indies in Antilles; South America to Peru
[WildflowerSearch map]
[BONAP county map]
Discussion

Species ca. 200 (4 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Blades 15-50 cm; areoles with included veinlets.
→ 2
1. Blades 5-15 cm; areoles rarely with included veinlets.
→ 3
2. Indusia peltate, attached centrally; pinnae cordate at base; rachises and costae glabrous abaxially.
T. heracleifolia
2. Indusia round-reniform, attached at sinus; pinnae oblique at base; rachises and costae puberulent abaxially.
T. incisa
3. Pinnae 1-8 pairs, often with proliferous buds in axils; petiole shorter than or rarely equaling blade, pubescent on both surfaces; indusia round-reniform, attached at sinus.
T. coriandrifolia
3. Pinnae absent or 1(-2) pair, without proliferous buds in axils; petiole 1-3 times length of blade, pubescent adaxially, lacking hairs abaxially; indusia peltate, attached centrally.
T. fimbriata
Source FNA vol. 2. Author: Robbin C. Moran. FNA vol. 2.
Parent taxa Dryopteridaceae Dryopteridaceae > Tectaria
Sibling taxa
T. coriandrifolia, T. fimbriata, T. incisa
Subordinate taxa
T. coriandrifolia, T. fimbriata, T. heracleifolia, T. incisa
Synonyms Aspidium heracleifolium
Name authority Cavanilles: Anales Hist. Nat. 1(2): 115. (1799) (Willdenow) L. Underwood: Bull. Torrey Bot. Club 33: 200. (1906)
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