Taraxacum ceratophorum |
Taraxacum hyparcticum |
|
---|---|---|
alpine dandelion, horned dandelion, pissenlit tuberculé |
high-arctic dandelion |
|
Habit | Plants (1–)6–50 cm; taproots branched. | Plants (3–)5–12(–30) cm; taproots occasionally branched. |
Stems | 1–10+, ascending to erect, ± purplish (at least proximally), densely villous (young) becoming glabrescent, sparsely villous to glabrate or glabrous proximally, ± densely villous distally. |
1–5, ascending to erect, pinkish to reddish or purplish, (barely exceeding foliage), glabrous or glabrate (rarely sparsely villous and villous distally). |
Leaves | ± 10, horizontal to patent, sometimes erect; sessile or petioles ± broadly winged (bases barely narrowed compared to blades); blades narrowly oblanceolate to linear-oblanceolate or linear-oblong (often ± runcinate), 4–30 × (0.4–)0.5–5 cm, bases cuneate to attenuate, margins lobed ± deeply to lacerate, irregularly to regularly, often toothed, merely denticulate, or subentire, lobes retrorse, straight or antrorse, deltate to triangular, acute to acuminate, teeth 0–1 on lobes, often more or mostly in sinuses, apices obtuse to sometimes acute, sometimes mucronate, faces glabrous or glabrate to very sparsely villous. |
fewer than 10, usually patent, rarely erect; petioles sometimes narrowly winged; blades oblanceolate (often ± runcinate), 2–12+ × (0.3–)0.5–1.2 cm, bases attenuate, margins lobed deeply and regularly to denticulate or subentire, lobes 5–6 pairs, straight to retrorse, triangular to deltate, teeth triangular 0–1 on lobes or 1–4 shallow pairs if subentire, acute to ± obtuse, apices obtuse to ± acute, faces glabrous. |
Involucres | dark green, sometimes ± glaucous, campanulate to ± hemispheric, (5–)8–19(–21) mm. |
dark green to bluish black or dark purplish green, narrowly campanulate, 15–30 mm. |
Florets | 40–85+; corollas yellow, drying cream to whitish (outer abaxially gray or purple-striped on drying), 10–22 × 1–2.8 mm. |
25–50; corollas cream-colored to white or pink-tinged distally, outer pinkish-striped abaxially, 15–20 × 1.2–3 mm. |
Phyllaries | (10–)12–14(–17) in 2 series, lanceolate to ovate-lanceolate (inner), 1.5–4.5 mm wide, margins scarious or not (outer), inner broadly scarious in proximal 1/2, apices usually horned, occasionally hornless, horns sometimes exceeding apices, tips white to purplish, scarious, erose. |
8–14 in 2 series, lanceolate to lance-ovate, 1.5–3.5 mm wide, margins narrowly (outer) to widely scarious in proximal 1/3 (inner), narrowly so distally, apices long-acuminate, sometimes callous, hornless (rarely very small horns), tips white-scarious, erose, rounded. |
Calyculi | of 12–16(–20), appressed to spreading, pale, ovate to elliptic or lance-ovate to lanceolate (sometimes thin) bractlets in 2–3 series, 5–12 × (0.9–)1.5–5 mm, margins hyaline, white or purplish, scarious, apices caudate to acuminate, ± strongly horned, callous, or occasionally some (rarely all) hornless, tips obtuse to rounded, scarious, erose. |
of 10–14, spreading to appressed (thinner than phyllaries), dark green, broadly ovate or ovate to oblong bractlets in 2–3 series, 6–10 × 3.4–5 mm, margins narrowly hyaline, scarious, apices abruptly acuminate to caudate, strongly horned, tip ± scarious, erose. |
Cypselae | olivaceous to olive brown, tan to olivaceous tan, brown to reddish brown, grayish brown or straw-colored, bodies oblanceoloid to obovoid, 2.5–4(–5) mm wide, cones conic or narrowly conic to broadly terete, 0.5–0.9 mm, beaks slender, 4.5–14 mm, ribs 5, large (bearing 10–15 narrower ones), faces proximally tuberculate or sometimes nearly smooth (usually with at least some tubercules) to muricate in distal 1/3–1/2, sometimes wholly muricate; pappi white to cream, 5–7.5(–8) mm. |
tan or straw-colored to brown (or reddish brown), sometimes grayish, bodies obovoid to oblanceoloid, 3–3.7(–4) mm, mostly broad, cones conic, 0.4–0.7 mm, beaks slender, 3.5–4.5 mm, ribs 10–13 (5–6 prominent), mostly broad, faces proximally tuberculate, muricate in distal 1/2; pappi whitish or sordid, 5–5.5(–7) mm. |
2n | = 16, [24], 32, 40, 48. |
= [24, 32, 40? some erroneous reports in literature under this name from Eurasia]. |
Taraxacum ceratophorum |
Taraxacum hyparcticum |
|
Phenology | Flowering spring–summer. | Flowering summer. |
Habitat | Wet to moist areas, calcareous or igneous rocks, gravel, sand, or clay, wet meadows, shores of streams, sandy or gravelly seashores, seepage slopes, early-melting snowbeds (south) | Dry, moderately drained areas in tundra, raised sand terraces, low center polygons on old surfaces, sandy, eroded knolls, marine/lacustrine deposits, rocky streambeds, dry slopes |
Elevation | 0–3000 m (0–9800 ft) | 20–1010 m (100–3300 ft) |
Distribution |
AK; CA; CO; MT; NM; NV; OR; UT; WA; WY; AB; BC; MB; NL; NT; NU; ON; QC; SK; YT; Greenland; Eurasia
|
AK; NT; NU; Greenland; Eurasia |
Discussion | Taraxacum ceratophorum is the most widespread native dandelion in North America, ranging from the low Arctic and boreal zone to the western Cordilleras, in the montane and alpine zones. This complex has been subdivided into many microspecies in North America, most of which appear unworthy of recognition. In the Quebec-Labrador Peninsula, Taraxacum ceratophorum grades continuously into what has been called T. hyperboreum. Inclusion of T. hyperboreum bridges the gap between typical T. ceratophorum and T. lacerum. Taraxacum lacerum stands out by its very lacerate leaves, but intermediates exist and it is impossible to draw a firm boundary. The lacerate Newfoundland form, T. longii, may be a spontaneous mutation within the range of T. ceratophorum. If T. lacerum were recognized, we would have to place T. longii within the former based on leaf morphology, though leaf orientation would be odd there (T. lacerum tends to have ascending leaves, and T. longii leaves that are flatter on the substrate). A more thorough morphometric and biosystematic study of this complex is warranted. Nonetheless, I have recognized two segregates (T. laurentianum, T. trigonolobum) that stand out from the continuum otherwise observed in the complex. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Taraxacum hyparcticum is sporadic in Arctic Eurasia (coastal Russia from Russian Far East west to Novaya Zemlya); it is mainly high-arctic. It is characterized by its small stature and large heads with white to yellowish cream ligules. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 19, p. 247. | FNA vol. 19, p. 249. |
Parent taxa | Asteraceae > tribe Cichorieae > Taraxacum | Asteraceae > tribe Cichorieae > Taraxacum |
Sibling taxa | ||
Synonyms | Leontodon ceratophorus, T. ambigens var. fultius, T. angulatum, T. arctogenum, T. brachyceras, T. carthamopsis, T. coverum, T. dumetorum, T. eriophorum, T. eurylepium, T. groenlandicum, T. hyperboreum, T. integratiforme, T. integratum, T. lacerum, T. lateritium, T. longii, T. mackenziense, T. malteanum, T. maurolepium, T. microcerum, T. ovinum, T. pellianum, T. pseudonorvegicum, T. umbriniforme, T. umbrinum | |
Name authority | (Ledebour) de Candolle: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 7: 146. (1838) | Dahlstedt: Ark. Bot. 4(8): 17, fig. 3. (1905) |
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