Syntrichia princeps |
Syntrichia ruralis |
|
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tortula moss |
sidewalk screw moss |
|
Stems | 5–20 mm. |
5–15 mm. |
Leaves | usually in distinct whorls, infolded, somewhat contorted, and weakly to strongly twisted around the stem when dry, wide-spreading to slightly recurved when moist, concave, spatulate, 2–4 × 1–1.5 mm; margins revolute in the proximal 1/2–3/4, entire; apices acute or sometimes truncate; costa excurrent into a long, serrate, hyaline awn (reddish at base), often strongly papillose abaxially and serrulate near the apex because of projecting cell ends, red; basal cells abruptly differentiated, long-rectangular, 45–80 × 20–30 µm, short-rectangular to quadrate at the margins; distal cells quadrate to hexagonal, 12–17 µm, slightly bulging, bearing 4–6 papillae per cell. |
clasping at base, infolded and twisted around the stem when dry, wide-spreading (in smaller forms) to squarrose-recurved when moist, lingulate-ovate, 1.5–3.5 × 0.75–1.25 mm, canaliculate to keeled; margins tightly revolute in the proximal 7/8 or more, entire; apices emarginate to acute; costa excurrent into a serrate (or occasionally only faintly serrulate), hyaline awn that is often red or sometimes broadly hyaline at base, weakly to strongly papillose on the abaxial surface and often serrate near the apex because of projecting cell ends, red-brown; basal cells abruptly differentiated, narrowly rectangular, 35–70(–90) × 11–18 µm, quadrate to narrowly rectangular at the margins; distal cells quadrate to polygonal, 8–12 µm, with 3–6 papillae per cell, bulging, somewhat obscure. |
Seta | red, 10–18 mm. |
red, 5–10 mm. |
Sexual condition | synoicous (apparently rarely dioicous). |
dioicous. |
Capsule | brownish red, 3–4 mm, slightly curved, with a distinct neck; operculum 1.5–2 mm, brown; peristome ca. 1.5 mm, the distal divisions twisted about 2 turns, red, the basal membrane white, 1/2–2/3 the total length. |
red-brown, 2–3.5 mm, straight, with an abrupt neck; operculum 1.25–1.75 mm, brown; peristome ca. 1.25 mm, the upper divisions twisted ca. 2 turns, red, the basal membrane white, about 1/3 the total length. |
Spores | 9–13 µm, papillose. |
11–15 µm, papillose. |
Specialized | asexual reproduction absent. |
asexual reproduction absent. |
Syntrichia princeps |
Syntrichia ruralis |
|
Habitat | Humus, soil, rock, tree bark | Dry to moist soil and rock |
Elevation | low to moderate elevations | low to high elevations |
Distribution |
AZ; CA; ID; MT; NV; OR; UT; WA; BC; Mexico; w South America; s South America; Europe; w Asia; Africa; Pacific Islands (Hawaii, New Zealand); Australia; Antarctica
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AK; AZ; CA; CO; ID; MA; ME; MI; MO; MT; NV; NY; OK; OR; SD; TX; UT; WA; WY; AB; BC; NF; NS; ON; Mexico; Eurasia; Greenland; s South America; s Africa |
Discussion | The synoicous condition of Syntrichia princeps is diagnostic if present, but otherwise one must rely on wider basal leaf cells, costal hydroids, and the stem central strand to separate this species from S. ruralis, S. papillosissima, and S. norvegica. The more acute leaves with cells generally smaller, and costa reddish and serrulate separate it from S. obtusissima. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Syntrichia ruralis generally has conspicuously squarrose-recurved leaves when wet, with margins recurved nearly to the apex, distal portions of the costa toothed abaxially because of projecting cell ends, and relatively small laminal cells. The decurrent, hyaline base of the awn sometimes used to distinguish S. ruraliformis is not reliable and can, on occasion, be found in S. princeps, S. papillosissima, and S. norvegica. Specific distinctions in the S. ruralis complex are subtle, for the most part, requiring cross sections of leaves and stems, and careful measurements. The leaves of S. princeps and S. obtusissima are narrowed near the middle, whereas those of S. papillosissima, S. norvegica, and S. ruralis are widest about one-third the way up from the base and then taper to the apex. The stem of S. princeps and S. obtusissima has a strong central strand of thinner-walled cells, the costa has a group of hydroids just abaxial to the guide cells, and the basal cells in the leaf are relatively wide. In S. papillosissima, S. norvegica, and S. ruralis, the stem lacks a central strand, the costa lacks hydroids, and the basal cells in the leaf are relatively narrow. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 27, p. 624. | FNA vol. 27, p. 627. |
Parent taxa | Pottiaceae > subfam. Pottioideae > Syntrichia | Pottiaceae > subfam. Pottioideae > Syntrichia |
Sibling taxa | ||
Synonyms | Tortula princeps | Barbula ruralis, S. ruraliformis, Tortula ruraliformis, Tortula ruralis |
Name authority | (De Notaris) Mitten: J. Proc. Linn. Soc., Bot., suppl. 1: 39. (1859) | (Hedwig) F. Weber & D. Mohr: Index Mus. Pl. Crypt., [2]. (1803) |
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