Syntrichia princeps |
Syntrichia papillosa |
|
---|---|---|
tortula moss |
||
Stems | 5–20 mm. |
1–4(–8) mm. |
Leaves | usually in distinct whorls, infolded, somewhat contorted, and weakly to strongly twisted around the stem when dry, wide-spreading to slightly recurved when moist, concave, spatulate, 2–4 × 1–1.5 mm; margins revolute in the proximal 1/2–3/4, entire; apices acute or sometimes truncate; costa excurrent into a long, serrate, hyaline awn (reddish at base), often strongly papillose abaxially and serrulate near the apex because of projecting cell ends, red; basal cells abruptly differentiated, long-rectangular, 45–80 × 20–30 µm, short-rectangular to quadrate at the margins; distal cells quadrate to hexagonal, 12–17 µm, slightly bulging, bearing 4–6 papillae per cell. |
incurved and slightly twisted when dry, erect to wide-spreading when moist, spatulate, (1.5–)2–3 × 0.75–1.25 mm; margins incurved when dry, plane to erect when moist, entire or occasionally serrulate near the apex; apices acute; costa excurrent into a short, smooth or serrulate, yellowish or hyaline awn 1/8–1/5 the leaf length, yellow or red, rounded and sharply papillose-serrate abaxially, smooth on the adaxial surface; basal cells gradually differentiated; distal cells isodiametric, rounded-hexagonal, 14–22 µm, papillae abaxial, single, simple, rarely forked near the costa, cells rather thick-walled, collenchymatous; cells elongate near leaf apex. |
Seta | red, 10–18 mm. |
|
Sexual condition | synoicous (apparently rarely dioicous). |
reportedly dioicous. |
Capsule | brownish red, 3–4 mm, slightly curved, with a distinct neck; operculum 1.5–2 mm, brown; peristome ca. 1.5 mm, the distal divisions twisted about 2 turns, red, the basal membrane white, 1/2–2/3 the total length. |
|
Spores | 9–13 µm, papillose. |
|
Specialized | asexual reproduction absent. |
asexual reproduction by gemmae borne on the adaxial surface of the costa, spherical or ovoid, 4–10-celled, brown when mature, smooth. |
Sporophytes | not known from the flora region. |
|
Syntrichia princeps |
Syntrichia papillosa |
|
Habitat | Humus, soil, rock, tree bark | Bark of trees or in rock crevices |
Elevation | low to moderate elevations | low to high elevations |
Distribution |
AZ; CA; ID; MT; NV; OR; UT; WA; BC; Mexico; w South America; s South America; Europe; w Asia; Africa; Pacific Islands (Hawaii, New Zealand); Australia; Antarctica
|
AZ; CA; CO; CT; GA; IL; MA; ME; MI; MO; NC; NJ; NM; NV; NY; OH; PA; TN; TX; UT; WA; NS; ON; Mexico; South America (Brazil, Colombia, Ecuador); Europe; s Africa; Atlantic Islands (Falkland Islands); Pacific Islands (New Zealand); Australia
|
Discussion | The synoicous condition of Syntrichia princeps is diagnostic if present, but otherwise one must rely on wider basal leaf cells, costal hydroids, and the stem central strand to separate this species from S. ruralis, S. papillosissima, and S. norvegica. The more acute leaves with cells generally smaller, and costa reddish and serrulate separate it from S. obtusissima. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The leaves of Syntrichia papillosa have unipapillose cells, with the papillae only on the abaxial surface, strongly papillose-serrate costae, and small, smooth, brown propagula borne on the adaxial surface of the costa. The plants superficially resemble S. laevipila in the field, but the position and nature of the propagula as well as the incurved leaf margins and roughened back of the costa distinguish S. papillosa, even with a hand lens. Sporophytes are known only from Australia, Tasmania, and New Zealand. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 27, p. 624. | FNA vol. 27, p. 620. |
Parent taxa | Pottiaceae > subfam. Pottioideae > Syntrichia | Pottiaceae > subfam. Pottioideae > Syntrichia |
Sibling taxa | ||
Synonyms | Tortula princeps | Tortula papillosa |
Name authority | (De Notaris) Mitten: J. Proc. Linn. Soc., Bot., suppl. 1: 39. (1859) | (Wilson) Juratska: Laubm.-Fl. Oestrr.-Ung., 141. (1882) |
Web links |
|