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tortula moss

Stems

5–20 mm.

(5–)10–20 mm.

Leaves

usually in distinct whorls, infolded, somewhat contorted, and weakly to strongly twisted around the stem when dry, wide-spreading to slightly recurved when moist, concave, spatulate, 2–4 × 1–1.5 mm;

margins revolute in the proximal 1/2–3/4, entire;

apices acute or sometimes truncate;

costa excurrent into a long, serrate, hyaline awn (reddish at base), often strongly papillose abaxially and serrulate near the apex because of projecting cell ends, red;

basal cells abruptly differentiated, long-rectangular, 45–80 × 20–30 µm, short-rectangular to quadrate at the margins;

distal cells quadrate to hexagonal, 12–17 µm, slightly bulging, bearing 4–6 papillae per cell.

clasping at base, infolded, and twisted around the stem when dry, wide-spreading to squarrose when moist, lingulate to spatulate, 3–4 × 1–1.5 mm, keeled;

margins revolute in the proximal 3/4–7/8, entire, often laxly undulate;

apices emarginate to retuse, occasionally truncate or acute;

costa excurrent into a toothed, hyaline awn (often brown at base) usually longer than the leaf, yellow or brown, often minutely papillose abaxially (but not serrate because of projecting cell ends);

basal cells abruptly differentiated, rectangular, 70–90(–100) × 22–40 µm, often with yellow walls, narrowly rectangular at the margins;

distal cells quadrate to polygonal, 15–23 µm, thin-walled, pellucid, bulging and bearing 4–7 papillae per cell.

Seta

red, 10–18 mm.

red-brown, 10–15 mm.

Sexual condition

synoicous (apparently rarely dioicous).

autoicous, although perigonia often lacking, apparently sometimes dioicous.

Capsule

brownish red, 3–4 mm, slightly curved, with a distinct neck;

operculum 1.5–2 mm, brown;

peristome ca. 1.5 mm, the distal divisions twisted about 2 turns, red, the basal membrane white, 1/2–2/3 the total length.

yellowish or reddish brown, 3–4.5 mm, straight or slightly curved, with an abrupt neck;

operculum 1.5–2 mm, reddish brown;

peristome ca. 1.5 mm, the distal divisions twisted ca. 2 turns, yellow or red, the basal membrane white, 1/3 the total length.

Spores

9–13 µm, papillose.

9–16 µm, papillose.

Specialized

asexual reproduction absent.

asexual reproduction absent.

Syntrichia princeps

Syntrichia obtusissima

Habitat Humus, soil, rock, tree bark Soil, rock, rarely the bases of trees
Elevation low to moderate elevations moderate to high elevations
Distribution
from FNA
AZ; CA; ID; MT; NV; OR; UT; WA; BC; Mexico; w South America; s South America; Europe; w Asia; Africa; Pacific Islands (Hawaii, New Zealand); Australia; Antarctica
[WildflowerSearch map]
from FNA
AZ; NM; TX; Mexico; South America (Bolivia, Peru)
Discussion

The synoicous condition of Syntrichia princeps is diagnostic if present, but otherwise one must rely on wider basal leaf cells, costal hydroids, and the stem central strand to separate this species from S. ruralis, S. papillosissima, and S. norvegica. The more acute leaves with cells generally smaller, and costa reddish and serrulate separate it from S. obtusissima.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Syntrichia obtusissima, with a distinctive Southwest and Mexican-Andean disjunction, has been much confused with S. ruralis, leading to many misidentified specimens in herbaria, but the large, pellucid leaf cells (about 15–23 µm) contrast with those of S. ruralis (about 8–12 µm), and the back of the costa lacks serrations from projecting cell ends. Furthermore, it has a clear central strand in the stem, and hydroids in the costa, neither of which is found in S. ruralis.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 27, p. 624. FNA vol. 27, p. 624.
Parent taxa Pottiaceae > subfam. Pottioideae > Syntrichia Pottiaceae > subfam. Pottioideae > Syntrichia
Sibling taxa
S. ammonsiana, S. amphidiacea, S. bartramii, S. cainii, S. caninervis, S. chisosa, S. fragilis, S. laevipila, S. latifolia, S. montana, S. norvegica, S. obtusissima, S. papillosa, S. papillosissima, S. ruralis, S. sinensis
S. ammonsiana, S. amphidiacea, S. bartramii, S. cainii, S. caninervis, S. chisosa, S. fragilis, S. laevipila, S. latifolia, S. montana, S. norvegica, S. papillosa, S. papillosissima, S. princeps, S. ruralis, S. sinensis
Synonyms Tortula princeps Barbula obtusissima, Tortula obtusissima
Name authority (De Notaris) Mitten: J. Proc. Linn. Soc., Bot., suppl. 1: 39. (1859) (Müller Hal.) R. H. Zander: Bull. Buffalo Soc. Nat. Sci. 32: 269. (1993)
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