Syntrichia princeps |
Syntrichia norvegica |
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tortula moss |
Norway screw moss, Norwegian tortula moss |
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Stems | 5–20 mm. |
8–25 mm. |
Leaves | usually in distinct whorls, infolded, somewhat contorted, and weakly to strongly twisted around the stem when dry, wide-spreading to slightly recurved when moist, concave, spatulate, 2–4 × 1–1.5 mm; margins revolute in the proximal 1/2–3/4, entire; apices acute or sometimes truncate; costa excurrent into a long, serrate, hyaline awn (reddish at base), often strongly papillose abaxially and serrulate near the apex because of projecting cell ends, red; basal cells abruptly differentiated, long-rectangular, 45–80 × 20–30 µm, short-rectangular to quadrate at the margins; distal cells quadrate to hexagonal, 12–17 µm, slightly bulging, bearing 4–6 papillae per cell. |
clasping at base, infolded and twisted around the stem when dry, squarrose-recurved when moist, lingulate-ovate, 2.5–3.5 × 1–1.2 mm, canaliculate to keeled; margins tightly revolute in the proximal 3/4; apices acute to acuminate; costa excurrent into a serrulate or sometimes serrate, hyaline awn that is often red at base or throughout but sometimes broadly hyaline at base, papillose abaxially and sometimes serrulate near the apex because of projecting cell ends, red-brown; basal cells abruptly differentiated, rectangular, 45–100 × 16–23 µm, quadrate to narrowly rectangular at the margins; distal cells quadrate, polygonal, or rectangular, 13–18 µm, with 3–6 papillae per cell, bulging, somewhat collenchymatous. |
Seta | red, 10–18 mm. |
brown, 15–20 mm. |
Sexual condition | synoicous (apparently rarely dioicous). |
dioicous. |
Capsule | brownish red, 3–4 mm, slightly curved, with a distinct neck; operculum 1.5–2 mm, brown; peristome ca. 1.5 mm, the distal divisions twisted about 2 turns, red, the basal membrane white, 1/2–2/3 the total length. |
red-brown, 3–4 mm, slightly curved, with an abrupt neck; operculum ca. 1.8 mm, brown; peristome ca. 1.8 mm, the upper divisions twisted ca. 2 turns, red, the basal membrane white, 1/3–1/2 the total length. |
Spores | 9–13 µm, papillose. |
11–15 µm, papillose. |
Specialized | asexual reproduction absent. |
asexual reproduction absent. |
Syntrichia princeps |
Syntrichia norvegica |
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Habitat | Humus, soil, rock, tree bark | Soil, rocks |
Elevation | low to moderate elevations | high elevations |
Distribution |
AZ; CA; ID; MT; NV; OR; UT; WA; BC; Mexico; w South America; s South America; Europe; w Asia; Africa; Pacific Islands (Hawaii, New Zealand); Australia; Antarctica
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AK; AZ; CA; CO; ID; MI; MT; NM; NV; OR; WA; AB; BC; ON; Mexico; Asia; Greenland; c Europe; n Europe; Africa (South Africa) |
Discussion | The synoicous condition of Syntrichia princeps is diagnostic if present, but otherwise one must rely on wider basal leaf cells, costal hydroids, and the stem central strand to separate this species from S. ruralis, S. papillosissima, and S. norvegica. The more acute leaves with cells generally smaller, and costa reddish and serrulate separate it from S. obtusissima. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Syntrichia norvegica can be distinguished from S. ruralis by its larger laminal cells, leaf margins less recurved distally, and apices consistently acute to acuminate. The awn is often partially to completely red, but the amount of color seems to vary with shade and is not considered definitive. Another potential recognition feature in the field is that the stereid band in the costa often disappears near the apex, making the normally reddish costa appear green just before the awn. Note that there are frequent neotenic forms of this species that lack an awn. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 27, p. 624. | FNA vol. 27, p. 626. |
Parent taxa | Pottiaceae > subfam. Pottioideae > Syntrichia | Pottiaceae > subfam. Pottioideae > Syntrichia |
Sibling taxa | ||
Synonyms | Tortula princeps | Tortula norvegica |
Name authority | (De Notaris) Mitten: J. Proc. Linn. Soc., Bot., suppl. 1: 39. (1859) | F. Weber: Arch. Syst. Naturgesch. 1(1): 130, plate 5, fig. 1. (1804) |
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