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tortula moss

Stems

5–20 mm.

2–10 mm.

Leaves

usually in distinct whorls, infolded, somewhat contorted, and weakly to strongly twisted around the stem when dry, wide-spreading to slightly recurved when moist, concave, spatulate, 2–4 × 1–1.5 mm;

margins revolute in the proximal 1/2–3/4, entire;

apices acute or sometimes truncate;

costa excurrent into a long, serrate, hyaline awn (reddish at base), often strongly papillose abaxially and serrulate near the apex because of projecting cell ends, red;

basal cells abruptly differentiated, long-rectangular, 45–80 × 20–30 µm, short-rectangular to quadrate at the margins;

distal cells quadrate to hexagonal, 12–17 µm, slightly bulging, bearing 4–6 papillae per cell.

infolded and twisted when dry, erect-spreading when moist, lingulate or spatulate, 1.5–3 × 0.5–1.5 mm;

margins revolute in the proximal 1/2, entire;

apices broadly acute or obtuse, sometimes emarginate;

costa excurrent into a short, serrate, hyaline awn, brown or red, often papillose abaxially and serrulate near the apex because of projecting cell ends;

basal cells abruptly differentiated, narrower toward the margins;

distal cells polygonal or quadrate, 8–12 µm, with 4–6 papillae per cell.

Seta

red, 10–18 mm.

red, 8–14 mm.

Sexual condition

synoicous (apparently rarely dioicous).

dioicous.

Capsule

brownish red, 3–4 mm, slightly curved, with a distinct neck;

operculum 1.5–2 mm, brown;

peristome ca. 1.5 mm, the distal divisions twisted about 2 turns, red, the basal membrane white, 1/2–2/3 the total length.

red, 2–3.5 mm, straight or slightly curved;

operculum 1.2–2 mm, red;

peristome 0.5–1.3 mm, the basal membrane to 1/2 the total length, red.

Spores

9–13 µm, papillose.

10–13 µm, papillose.

Specialized

asexual reproduction absent.

asexual reproduction absent.

Syntrichia princeps

Syntrichia montana

Habitat Humus, soil, rock, tree bark Widespread on soil and rock, occasionally on tree bark
Elevation low to moderate elevations moderate to high elevations
Distribution
from FNA
AZ; CA; ID; MT; NV; OR; UT; WA; BC; Mexico; w South America; s South America; Europe; w Asia; Africa; Pacific Islands (Hawaii, New Zealand); Australia; Antarctica
[WildflowerSearch map]
from FNA
AZ; CA; ID; ME; MT; NM; NV; NY; OR; WA; AB; BC; Europe; Asia
Discussion

The synoicous condition of Syntrichia princeps is diagnostic if present, but otherwise one must rely on wider basal leaf cells, costal hydroids, and the stem central strand to separate this species from S. ruralis, S. papillosissima, and S. norvegica. The more acute leaves with cells generally smaller, and costa reddish and serrulate separate it from S. obtusissima.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

This treatment follows M. T. Gallego et al. (2006) in employing Syntrichia montana to encompass what has been called S. intermedia or S. ruralis var. crinita in North America. This is, however, a tentative decision as there remains an incompletely understood complex of small, dioicous plants with hydroids in the costa, and some currently undescribed taxa may yet need to be recognized. One recently described species that falls in this complex is S. rupicola B. H. Allen. Further field sampling, molecular characterization, and culture studies are necessary before these taxa are fully understood.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 27, p. 624. FNA vol. 27, p. 624.
Parent taxa Pottiaceae > subfam. Pottioideae > Syntrichia Pottiaceae > subfam. Pottioideae > Syntrichia
Sibling taxa
S. ammonsiana, S. amphidiacea, S. bartramii, S. cainii, S. caninervis, S. chisosa, S. fragilis, S. laevipila, S. latifolia, S. montana, S. norvegica, S. obtusissima, S. papillosa, S. papillosissima, S. ruralis, S. sinensis
S. ammonsiana, S. amphidiacea, S. bartramii, S. cainii, S. caninervis, S. chisosa, S. fragilis, S. laevipila, S. latifolia, S. norvegica, S. obtusissima, S. papillosa, S. papillosissima, S. princeps, S. ruralis, S. sinensis
Synonyms Tortula princeps S. intermedia, Tortula intermedia, Tortula ruralis var. crinita
Name authority (De Notaris) Mitten: J. Proc. Linn. Soc., Bot., suppl. 1: 39. (1859) Nees: Flora 2: 301. (1819)
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