Syntrichia princeps |
Syntrichia laevipila |
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tortula moss |
twisted oak moss, twisted screw moss |
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Stems | 5–20 mm. |
1–3(–5) mm. |
Leaves | usually in distinct whorls, infolded, somewhat contorted, and weakly to strongly twisted around the stem when dry, wide-spreading to slightly recurved when moist, concave, spatulate, 2–4 × 1–1.5 mm; margins revolute in the proximal 1/2–3/4, entire; apices acute or sometimes truncate; costa excurrent into a long, serrate, hyaline awn (reddish at base), often strongly papillose abaxially and serrulate near the apex because of projecting cell ends, red; basal cells abruptly differentiated, long-rectangular, 45–80 × 20–30 µm, short-rectangular to quadrate at the margins; distal cells quadrate to hexagonal, 12–17 µm, slightly bulging, bearing 4–6 papillae per cell. |
infolded and somewhat twisted when dry, erect-spreading when moist, spatulate, 1–1.75 × 0.5–0.75 mm; margins plane (or somewhat erect), crenulate; apices obtuse, blunt, or emarginate; costa excurrent into a conspicuously tapered, smooth, yellowish or hyaline awn 1/5–1/2 the length of the leaf, yellow or red, rounded and smooth on abaxial surface; basal cells rather abruptly differentiated, often with thickened cross-walls, usually extending slightly farther up the margin than near the costa; distal cells quadrate to hexagonal, 10–15 µm, bulging, with about 4 papillae per cell, the outermost row of cells less papillose. |
Seta | red, 10–18 mm. |
red, 5–15 mm. |
Sexual condition | synoicous (apparently rarely dioicous). |
dioicous or autoicous. |
Capsule | brownish red, 3–4 mm, slightly curved, with a distinct neck; operculum 1.5–2 mm, brown; peristome ca. 1.5 mm, the distal divisions twisted about 2 turns, red, the basal membrane white, 1/2–2/3 the total length. |
red, 3–5 mm, straight, with a distinct neck; operculum 1–2 mm, red; peristome 0.5–1 mm, basal membrane 1/2 the total length, red. |
Spores | 9–13 µm, papillose. |
10–16 µm, papillose. |
Specialized | asexual reproduction absent. |
asexual reproduction as propagula borne at the tip of the stem and in axils of distal leaves, fusiform, green, ecostate, papillose, with an elongate terminal cell, occasionally as several cells. |
Syntrichia princeps |
Syntrichia laevipila |
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Habitat | Humus, soil, rock, tree bark | Occasional on bark of trees, rarely on rock |
Elevation | low to moderate elevations | low to high elevations |
Distribution |
AZ; CA; ID; MT; NV; OR; UT; WA; BC; Mexico; w South America; s South America; Europe; w Asia; Africa; Pacific Islands (Hawaii, New Zealand); Australia; Antarctica
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AL; AR; AZ; CA; CO; CT; GA; KS; KY; LA; MD; MO; NC; NE; NM; NV; OH; OK; OR; SC; SD; TN; TX; VA; WA; WV; BC; Mexico; South America (Chile); s Europe; e Asia (Japan); s Africa; Pacific Islands (New Zealand); Australia
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Discussion | The synoicous condition of Syntrichia princeps is diagnostic if present, but otherwise one must rely on wider basal leaf cells, costal hydroids, and the stem central strand to separate this species from S. ruralis, S. papillosissima, and S. norvegica. The more acute leaves with cells generally smaller, and costa reddish and serrulate separate it from S. obtusissima. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
This treatment follows M. T. Gallego et al. (2006) in including Syntrichia pagorum with forms of S. laevipila reported in the literature from North America, but this is tentative pending a careful reexamination of this complex of small, corticolous plants. These plants might be confused superfically with S. papillosa, but the leaf margins are plane, the abaxial surface of the costa is smooth and shiny, the propagula are leaf-like, and the leaf cells are pluripapillose in S. laevipila. The other taxa with leaf-like propagula, S. chisosa and S. ammonsiana, are never corticolous. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 27, p. 624. | FNA vol. 27, p. 621. |
Parent taxa | Pottiaceae > subfam. Pottioideae > Syntrichia | Pottiaceae > subfam. Pottioideae > Syntrichia |
Sibling taxa | ||
Synonyms | Tortula princeps | S. pagorum, Tortula laevipila var. meridionalis, Tortula pagorum |
Name authority | (De Notaris) Mitten: J. Proc. Linn. Soc., Bot., suppl. 1: 39. (1859) | Bridel: Muscol. Recent., suppl. 4: 98. (1818) |
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