The green links below add additional plants to the comparison table. Blue links lead to other Web sites.
enable glossary links

tortula moss

Stems

5–20 mm.

(5–)10–25 mm.

Leaves

usually in distinct whorls, infolded, somewhat contorted, and weakly to strongly twisted around the stem when dry, wide-spreading to slightly recurved when moist, concave, spatulate, 2–4 × 1–1.5 mm;

margins revolute in the proximal 1/2–3/4, entire;

apices acute or sometimes truncate;

costa excurrent into a long, serrate, hyaline awn (reddish at base), often strongly papillose abaxially and serrulate near the apex because of projecting cell ends, red;

basal cells abruptly differentiated, long-rectangular, 45–80 × 20–30 µm, short-rectangular to quadrate at the margins;

distal cells quadrate to hexagonal, 12–17 µm, slightly bulging, bearing 4–6 papillae per cell.

crowded, longitudinally folded and spirally twisted around the stem (but little crisped) when dry, wide-spreading when moist, oblong-lingulate to spatulate, 2–3.5 × 0.75–1 mm, fragile, with sheets of cells breaking off, sometimes along lines of weakness;

margins revolute before to as much as 1/2 the leaf length, to plane, entire or crenulate;

apices truncate to acute;

costa percurrent or slightly excurrent as a mucro 1–3(–5) cells long, yellow or red, finely to strongly papillose abaxially, abruptly tapered;

basal cells abruptly differentiated, becoming rather abruptly short-rectangular near the margins;

distal cells quadrate-hexagonal, 9–15 µm, bulging, with 3–8 papillae per cell, with moderately thick walls, not particularly collenchymatous;

distal marginal cells not or weakly to strongly differentiated as a border of brownish, thicker-walled cells in about 2–3 rows.

Seta

red, 10–18 mm.

red, 10–15 mm.

Sexual condition

synoicous (apparently rarely dioicous).

dioicous.

Capsule

brownish red, 3–4 mm, slightly curved, with a distinct neck;

operculum 1.5–2 mm, brown;

peristome ca. 1.5 mm, the distal divisions twisted about 2 turns, red, the basal membrane white, 1/2–2/3 the total length.

red, (1.5–)2–4 mm, slightly curved, rather gradually tapered to the seta or with a distinct neck;

operculum 1–1.5 mm, red;

peristome 1–1.5 mm, teeth twisted about 1/2 turn, the distal divisions yellow, the basal membrane pale, about 1/4 the total length.

Spores

9–13 µm, papillose.

13–20 µm, densely papillose.

Specialized

asexual reproduction absent.

asexual reproduction absent.

Syntrichia princeps

Syntrichia fragilis

Habitat Humus, soil, rock, tree bark Bark of trees, rock (usually calcareous), occasionally soil
Elevation low to moderate elevations low to high elevations
Distribution
from FNA
AZ; CA; ID; MT; NV; OR; UT; WA; BC; Mexico; w South America; s South America; Europe; w Asia; Africa; Pacific Islands (Hawaii, New Zealand); Australia; Antarctica
[WildflowerSearch map]
from FNA
AZ; NC; NM; OK; TN; TX; VA; WV; HI; Mexico; Central America; South America; Europe; Asia (China, Iran)
Discussion

The synoicous condition of Syntrichia princeps is diagnostic if present, but otherwise one must rely on wider basal leaf cells, costal hydroids, and the stem central strand to separate this species from S. ruralis, S. papillosissima, and S. norvegica. The more acute leaves with cells generally smaller, and costa reddish and serrulate separate it from S. obtusissima.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Syntrichia fragilis shows considerable variability in size and leaf shape (apparently correlated with habitat conditions). In some small, corticolous plants, extreme fragmentation is evident, with an abnormal development of the lamina beyond the usual outline of the leaf. In such plants, several regions of each leaf seem to remain meristematic and produce scalloped or ruffled extensions that eventually break off. Fragility of leaves should not be considered an infallibly diagnostic character, however, since some plants of S. fragilis have quite firm leaves, and the leaves of some other species, particularly when old, may become broken.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 27, p. 624. FNA vol. 27, p. 623.
Parent taxa Pottiaceae > subfam. Pottioideae > Syntrichia Pottiaceae > subfam. Pottioideae > Syntrichia
Sibling taxa
S. ammonsiana, S. amphidiacea, S. bartramii, S. cainii, S. caninervis, S. chisosa, S. fragilis, S. laevipila, S. latifolia, S. montana, S. norvegica, S. obtusissima, S. papillosa, S. papillosissima, S. ruralis, S. sinensis
S. ammonsiana, S. amphidiacea, S. bartramii, S. cainii, S. caninervis, S. chisosa, S. laevipila, S. latifolia, S. montana, S. norvegica, S. obtusissima, S. papillosa, S. papillosissima, S. princeps, S. ruralis, S. sinensis
Synonyms Tortula princeps Tortula fragilis
Name authority (De Notaris) Mitten: J. Proc. Linn. Soc., Bot., suppl. 1: 39. (1859) (Taylor) Ochyra: Fragm. Florist. Geobot. 37: 212. (1992)
Web links