Syntrichia princeps |
Syntrichia chisosa |
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tortula moss |
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Stems | 5–20 mm. |
2–6 mm. |
Leaves | usually in distinct whorls, infolded, somewhat contorted, and weakly to strongly twisted around the stem when dry, wide-spreading to slightly recurved when moist, concave, spatulate, 2–4 × 1–1.5 mm; margins revolute in the proximal 1/2–3/4, entire; apices acute or sometimes truncate; costa excurrent into a long, serrate, hyaline awn (reddish at base), often strongly papillose abaxially and serrulate near the apex because of projecting cell ends, red; basal cells abruptly differentiated, long-rectangular, 45–80 × 20–30 µm, short-rectangular to quadrate at the margins; distal cells quadrate to hexagonal, 12–17 µm, slightly bulging, bearing 4–6 papillae per cell. |
infolded and spirally twisted around the stem when dry, wide-spreading to squarrose-recurved when moist, spatulate, 1.5–2.5 × 0.6–0.8(–1) mm, distally 2-stratose, occasionally with 1-stratose patches; margins plane, entire; apices acute to truncate; costa excurrent into a short, 0.02–0.1 mm mucro, red, rounded and sharply serrate with hyaline teeth abaxially; basal cells abruptly differentiated, those near the margin smaller, short-rectangular; distal cells hexagonal to irregularly polygonal, or isodiametric, 8–12(–13) µm, densely papillose with 3–5 papillae per cell, bulging, moderately thick-walled, not collenchymatous, obscure. |
Seta | red, 10–18 mm. |
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Sexual condition | synoicous (apparently rarely dioicous). |
dioicous (perigonia and sporophytes unknown). |
Capsule | brownish red, 3–4 mm, slightly curved, with a distinct neck; operculum 1.5–2 mm, brown; peristome ca. 1.5 mm, the distal divisions twisted about 2 turns, red, the basal membrane white, 1/2–2/3 the total length. |
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Spores | 9–13 µm, papillose. |
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Specialized | asexual reproduction absent. |
asexual reproduction by propagula borne on stalks in axils of distal leaves, leaf-like, squarrose-recurved, 0.25–0.45 mm, green, papillose, costate, with an apiculus of 1-several elongate, smooth cells. |
Syntrichia princeps |
Syntrichia chisosa |
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Habitat | Humus, soil, rock, tree bark | Locally common in desert scrub, thin soil in crevices of rocks, soil around bases of shrubs and boulders |
Elevation | low to moderate elevations | moderate to high elevations (about 1500-2700 m) (moderate to high elevations (about 4900-8900 ft)) |
Distribution |
AZ; CA; ID; MT; NV; OR; UT; WA; BC; Mexico; w South America; s South America; Europe; w Asia; Africa; Pacific Islands (Hawaii, New Zealand); Australia; Antarctica
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NM; OK; TX; Mexico; s Africa |
Discussion | The synoicous condition of Syntrichia princeps is diagnostic if present, but otherwise one must rely on wider basal leaf cells, costal hydroids, and the stem central strand to separate this species from S. ruralis, S. papillosissima, and S. norvegica. The more acute leaves with cells generally smaller, and costa reddish and serrulate separate it from S. obtusissima. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Distinguishing features of Syntrichia chisosa include the 2-stratose laminae, strongly serrate abaxial surface of the costa, plane leaf margins, mucronate apex, and costate, foliose propagula. In the northern part of its range, the species frequently occurs with S. bartramii (see comments thereunder). Syntrichia chisosa also occurs with S. fragilis, and the two may be confused because of a superficially similar habit, color, and dry leaf stance, but S. chisosa can be distinguished with a handlens by non-fragile leaves, a fuzzy appearance of the back of the costa, and usually abundant propagula, and, under a compound microscope, by smaller, 2-stratose, and relatively obscure laminal cells. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 27, p. 624. | FNA vol. 27, p. 622. |
Parent taxa | Pottiaceae > subfam. Pottioideae > Syntrichia | Pottiaceae > subfam. Pottioideae > Syntrichia |
Sibling taxa | ||
Synonyms | Tortula princeps | Tortula chisosa |
Name authority | (De Notaris) Mitten: J. Proc. Linn. Soc., Bot., suppl. 1: 39. (1859) | (Magill Delgadillo & L. R. Stark) R. H. Zander: Bull. Buffalo Soc. Nat. Sci. 32: 269. (1993) |
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