Syntrichia princeps |
Syntrichia caninervis |
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tortula moss |
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Stems | 5–20 mm. |
3–20 mm. |
Leaves | usually in distinct whorls, infolded, somewhat contorted, and weakly to strongly twisted around the stem when dry, wide-spreading to slightly recurved when moist, concave, spatulate, 2–4 × 1–1.5 mm; margins revolute in the proximal 1/2–3/4, entire; apices acute or sometimes truncate; costa excurrent into a long, serrate, hyaline awn (reddish at base), often strongly papillose abaxially and serrulate near the apex because of projecting cell ends, red; basal cells abruptly differentiated, long-rectangular, 45–80 × 20–30 µm, short-rectangular to quadrate at the margins; distal cells quadrate to hexagonal, 12–17 µm, slightly bulging, bearing 4–6 papillae per cell. |
infolded and imbricate, not to weakly twisted around stem when dry, erect-spreading when moist, ovate-spatulate, 1–2.5 × 0.6–1.2 mm, 2-stratose or thicker, occasionally with scattered 1-stratose patches; margins revolute for the entire length of leaf, entire; apices acute to acuminate; costa excurrent into a serrate, hyaline awn that is often broadly hyaline at base, brown or blackish, strongly papillose; basal cells abruptly differentiated, narrower toward the margins; distal cells rounded, polygonal, or quadrate, 8–13 µm, with 4–6 low papillae per cell. |
Seta | red, 10–18 mm. |
6–15 mm, brown. |
Sexual condition | synoicous (apparently rarely dioicous). |
dioicous. |
Capsule | brownish red, 3–4 mm, slightly curved, with a distinct neck; operculum 1.5–2 mm, brown; peristome ca. 1.5 mm, the distal divisions twisted about 2 turns, red, the basal membrane white, 1/2–2/3 the total length. |
red, 2.5–3.2 mm, straight or slightly curved, with a distinct neck; operculum 1.5–2mm, red; peristome 0.7–1 mm, the basal membrane about 1/2 the total length. |
Spores | 9–13 µm, papillose. |
7–8 µm, lightly papillose. |
Specialized | asexual reproduction absent. |
asexual reproduction absent. |
Syntrichia princeps |
Syntrichia caninervis |
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Habitat | Humus, soil, rock, tree bark | Soil, deserts and steppe, often forming extensive carpets |
Elevation | low to moderate elevations | moderate to high elevations |
Distribution |
AZ; CA; ID; MT; NV; OR; UT; WA; BC; Mexico; w South America; s South America; Europe; w Asia; Africa; Pacific Islands (Hawaii, New Zealand); Australia; Antarctica
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AZ; CA; CO; ID; MT; NM; NV; OR; UT; WA; WY; AB; BC; s Asia; sw Asia (Afghanistan, Iran, Lebanon, Russia, Turkey); n Africa |
Discussion | The synoicous condition of Syntrichia princeps is diagnostic if present, but otherwise one must rely on wider basal leaf cells, costal hydroids, and the stem central strand to separate this species from S. ruralis, S. papillosissima, and S. norvegica. The more acute leaves with cells generally smaller, and costa reddish and serrulate separate it from S. obtusissima. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Syntrichia caninervis is most common in the colder deserts and steppes of the flora area, particularly in the Mojave and Great Basin deserts and the Columbia Basin. It can be confused in the field with S. ruralis, but good field distinctions for S. caninervis include the blackish or olive green color, the imbricate, weakly twisted leaf stance when dry, and the back of the costa showing no trace of red and often having a frosty appearance because of the stellate papillae. Microscopically, S. caninervis is unique with its combination of 2-stratose laminae, non-bulging cell surfaces, and costal cross-sections with sub-stereid cells. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 27, p. 624. | FNA vol. 27, p. 625. |
Parent taxa | Pottiaceae > subfam. Pottioideae > Syntrichia | Pottiaceae > subfam. Pottioideae > Syntrichia |
Sibling taxa | ||
Synonyms | Tortula princeps | Tortula bistratosa, Tortula caninervis |
Name authority | (De Notaris) Mitten: J. Proc. Linn. Soc., Bot., suppl. 1: 39. (1859) | Mitten: J. Proc. Linn. Soc., Bot., suppl. 1: 39. (1859) |
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