Syntrichia princeps |
Syntrichia bartramii |
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tortula moss |
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Stems | 5–20 mm. |
2–10 mm. |
Leaves | usually in distinct whorls, infolded, somewhat contorted, and weakly to strongly twisted around the stem when dry, wide-spreading to slightly recurved when moist, concave, spatulate, 2–4 × 1–1.5 mm; margins revolute in the proximal 1/2–3/4, entire; apices acute or sometimes truncate; costa excurrent into a long, serrate, hyaline awn (reddish at base), often strongly papillose abaxially and serrulate near the apex because of projecting cell ends, red; basal cells abruptly differentiated, long-rectangular, 45–80 × 20–30 µm, short-rectangular to quadrate at the margins; distal cells quadrate to hexagonal, 12–17 µm, slightly bulging, bearing 4–6 papillae per cell. |
infolded and spirally twisted around the stem when dry, wide-spreading when moist, lingulate to spatulate, 1.25–2 × 0.5–0.75 mm; margins plane, entire, except for papillose crenulations, not bordered; apices acute to truncate or occasionally emarginate; costa excurrent into a conspicuously tapered, serrate awn 0.1–0.6 mm, red or yellow, sparsely to densely spinulose abaxially; basal cells abruptly differentiated, with somewhat thickened cross walls, those at the margins narrower; distal cells irregularly polygonal, isodiametric, 9–13 µm, obscure, bulging and densely papillose, with 4–6 papillae per cell, moderately thick-walled and not collenchymatous. |
Seta | red, 10–18 mm. |
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Sexual condition | synoicous (apparently rarely dioicous). |
dioicous (perigonia and sporophytes unknown). |
Capsule | brownish red, 3–4 mm, slightly curved, with a distinct neck; operculum 1.5–2 mm, brown; peristome ca. 1.5 mm, the distal divisions twisted about 2 turns, red, the basal membrane white, 1/2–2/3 the total length. |
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Spores | 9–13 µm, papillose. |
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Specialized | asexual reproduction absent. |
asexual reproduction absent. |
Syntrichia princeps |
Syntrichia bartramii |
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Habitat | Humus, soil, rock, tree bark | Dry soil and rocks |
Elevation | low to moderate elevations | moderate to high elevations |
Distribution |
AZ; CA; ID; MT; NV; OR; UT; WA; BC; Mexico; w South America; s South America; Europe; w Asia; Africa; Pacific Islands (Hawaii, New Zealand); Australia; Antarctica
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AZ; CA; CO; ID; NM; TX; Mexico (Baja California) |
Discussion | The synoicous condition of Syntrichia princeps is diagnostic if present, but otherwise one must rely on wider basal leaf cells, costal hydroids, and the stem central strand to separate this species from S. ruralis, S. papillosissima, and S. norvegica. The more acute leaves with cells generally smaller, and costa reddish and serrulate separate it from S. obtusissima. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Diagnostic features of the rare Syntrichia bartramii include plane leaf margins and the costa excurrent into a tapered, serrate awn. In contrast with those of S. fragilis, the leaves of S. bartramii are not fragile and have awns, somewhat smaller, more obscure distal cells, less well-defined groups of basal cells, and plane margins. Without propagula, plants of S. laevipila can be separated from S. bartramii by the abaxially smooth costae and smooth awns, and also by larger and less obscure leaf cells. It should be noted that leaves of S. bartramii sometimes show 2-stratose patches (one or a few cells in width). In contrast, S. chisosa has leaves that are almost completely 2-stratose distally, less twisted when dry, and merely mucronate at the apex. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 27, p. 624. | FNA vol. 27, p. 621. |
Parent taxa | Pottiaceae > subfam. Pottioideae > Syntrichia | Pottiaceae > subfam. Pottioideae > Syntrichia |
Sibling taxa | ||
Synonyms | Tortula princeps | Tortula bartramii |
Name authority | (De Notaris) Mitten: J. Proc. Linn. Soc., Bot., suppl. 1: 39. (1859) | (Steere) R. H. Zander: Bull. Buffalo Soc. Nat. Sci. 32: 267. (1993) |
Web links |
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