Syntrichia princeps |
Syntrichia papillosissima |
|
---|---|---|
tortula moss |
|
|
Stems | 5–20 mm. |
10–25 mm. |
Leaves | usually in distinct whorls, infolded, somewhat contorted, and weakly to strongly twisted around the stem when dry, wide-spreading to slightly recurved when moist, concave, spatulate, 2–4 × 1–1.5 mm; margins revolute in the proximal 1/2–3/4, entire; apices acute or sometimes truncate; costa excurrent into a long, serrate, hyaline awn (reddish at base), often strongly papillose abaxially and serrulate near the apex because of projecting cell ends, red; basal cells abruptly differentiated, long-rectangular, 45–80 × 20–30 µm, short-rectangular to quadrate at the margins; distal cells quadrate to hexagonal, 12–17 µm, slightly bulging, bearing 4–6 papillae per cell. |
clasping at base, infolded and twisted around the stem when dry, squarrose-recurved when moist, lingulate-ovate, 2.5–4 × 1–1.6 mm, canaliculate to keeled; margins tightly revolute in the proximal 3/4–7/8, entire; apices obtuse to acute; costa excurrent into a serrate, hyaline awn that is often brown, sometimes broadly hyaline at base, strongly papillose abaxially and serrate because of projecting cell ends, yellow-brown; basal cells abruptly differentiated, rectangular, 45–90 × 15–23 µm, quadrate to narrowly rectangular at the margins; distal cells quadrate to polygonal, 11–18 µm, with tall, bulging mammillae, bearing 1–2 papillae per cell, thick-walled and sometimes collenchymatous. |
Seta | red, 10–18 mm. |
brown, 12–18 mm. |
Sexual condition | synoicous (apparently rarely dioicous). |
dioicous. |
Capsule | brownish red, 3–4 mm, slightly curved, with a distinct neck; operculum 1.5–2 mm, brown; peristome ca. 1.5 mm, the distal divisions twisted about 2 turns, red, the basal membrane white, 1/2–2/3 the total length. |
brown, 3–5 mm, curved, with an abrupt neck; operculum ca. 2 mm; peristome ca. 1.8 mm, the upper divisions twisted ca. 2 turns, yellow-brown, the basal membrane white, ca. 1/2 the total length. |
Spores | 9–13 µm, papillose. |
10–14 µm, papillose. |
Specialized | asexual reproduction absent. |
asexual reproduction absent. |
Syntrichia princeps |
Syntrichia papillosissima |
|
Habitat | Humus, soil, rock, tree bark | Dry soil, rock |
Elevation | low to moderate elevations | moderate to high elevations |
Distribution |
AZ; CA; ID; MT; NV; OR; UT; WA; BC; Mexico; w South America; s South America; Europe; w Asia; Africa; Pacific Islands (Hawaii, New Zealand); Australia; Antarctica
|
AZ; CA; CO; ID; MT; NM; NV; OR; UT; WA; BC; Mexico (Coahuila, Nuevo León); s Europe; c Asia |
Discussion | The synoicous condition of Syntrichia princeps is diagnostic if present, but otherwise one must rely on wider basal leaf cells, costal hydroids, and the stem central strand to separate this species from S. ruralis, S. papillosissima, and S. norvegica. The more acute leaves with cells generally smaller, and costa reddish and serrulate separate it from S. obtusissima. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Syntrichia papillosissima is primarily a species of the Great Basin Desert north into the shrub-steppe ecosystems of the Columbia Basin, where it often occurs as a co-dominant with S. ruralis and S. caninervis. It is similar to a robust S. ruralis, differing most conspicuously in the extremely tall mammillae on the distal leaf cells, each crowned by only one or two papillae, unlike the shorter bulging cell surface bearing four or five papillae characterizing other species in the S. ruralis complex. Syntrichia papillosissima also has larger distal laminal cells, which are more pellucid than those of S. ruralis. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 27, p. 624. | FNA vol. 27, p. 626. |
Parent taxa | Pottiaceae > subfam. Pottioideae > Syntrichia | Pottiaceae > subfam. Pottioideae > Syntrichia |
Sibling taxa | ||
Synonyms | Tortula princeps | Barbula papillosissima, Barbula ruralis var. hirsuta, Tortula papillosissima, Tortula ruralis var. hirsuta |
Name authority | (De Notaris) Mitten: J. Proc. Linn. Soc., Bot., suppl. 1: 39. (1859) | (Coppey) Loeske: Hedwigia 49: 44. (1910) |
Web links |
|
|