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tortula moss

wideleaf tortula moss

Stems

5–20 mm.

4–12 mm.

Leaves

usually in distinct whorls, infolded, somewhat contorted, and weakly to strongly twisted around the stem when dry, wide-spreading to slightly recurved when moist, concave, spatulate, 2–4 × 1–1.5 mm;

margins revolute in the proximal 1/2–3/4, entire;

apices acute or sometimes truncate;

costa excurrent into a long, serrate, hyaline awn (reddish at base), often strongly papillose abaxially and serrulate near the apex because of projecting cell ends, red;

basal cells abruptly differentiated, long-rectangular, 45–80 × 20–30 µm, short-rectangular to quadrate at the margins;

distal cells quadrate to hexagonal, 12–17 µm, slightly bulging, bearing 4–6 papillae per cell.

infolded, twisted, and slightly contorted when dry, erect-spreading when moist, narrowly or broadly spatulate, 1.8–2.8 × 0.6–0.8 mm;

margins revolute in the proximal 1/2, entire;

apices obtuse;

costa ending a few cells before the apex or percurrent, brown, smooth;

basal cells abruptly differentiated, narrower toward the margins;

distal cells rounded, polygonal, or quadrate, 12–18 µm, with 4–6 papillae per cell.

Seta

red, 10–18 mm.

red, 6 mm.

Sexual condition

synoicous (apparently rarely dioicous).

dioicous.

Capsule

brownish red, 3–4 mm, slightly curved, with a distinct neck;

operculum 1.5–2 mm, brown;

peristome ca. 1.5 mm, the distal divisions twisted about 2 turns, red, the basal membrane white, 1/2–2/3 the total length.

red, 3 mm, straight, with a distinct neck;

operculum 1.5–2mm, red;

peristome 1 mm, basal membrane 1/2–2/3 the total length, red.

Spores

9–13 µm, papillose.

10–15 µm, papillose.

Specialized

asexual reproduction absent.

asexual reproduction by gemmae on the adaxial or occasionally the abaxial leaf surface of leaves, ovoid and rounded, 25–45 µm in diameter, green to brown, multicellular.

Syntrichia princeps

Syntrichia latifolia

Habitat Humus, soil, rock, tree bark Bark of trees, rarely on rocks
Elevation low to moderate elevations low to high elevations
Distribution
from FNA
AZ; CA; ID; MT; NV; OR; UT; WA; BC; Mexico; w South America; s South America; Europe; w Asia; Africa; Pacific Islands (Hawaii, New Zealand); Australia; Antarctica
[WildflowerSearch map]
from FNA
AZ; CA; OR; WA; BC; Europe
Discussion

The synoicous condition of Syntrichia princeps is diagnostic if present, but otherwise one must rely on wider basal leaf cells, costal hydroids, and the stem central strand to separate this species from S. ruralis, S. papillosissima, and S. norvegica. The more acute leaves with cells generally smaller, and costa reddish and serrulate separate it from S. obtusissima.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 27, p. 624. FNA vol. 27, p. 620.
Parent taxa Pottiaceae > subfam. Pottioideae > Syntrichia Pottiaceae > subfam. Pottioideae > Syntrichia
Sibling taxa
S. ammonsiana, S. amphidiacea, S. bartramii, S. cainii, S. caninervis, S. chisosa, S. fragilis, S. laevipila, S. latifolia, S. montana, S. norvegica, S. obtusissima, S. papillosa, S. papillosissima, S. ruralis, S. sinensis
S. ammonsiana, S. amphidiacea, S. bartramii, S. cainii, S. caninervis, S. chisosa, S. fragilis, S. laevipila, S. montana, S. norvegica, S. obtusissima, S. papillosa, S. papillosissima, S. princeps, S. ruralis, S. sinensis
Synonyms Tortula princeps Tortula latifolia
Name authority (De Notaris) Mitten: J. Proc. Linn. Soc., Bot., suppl. 1: 39. (1859) (Bruch ex Hartman) Huebener: Muscol. Germ., 342. (1833)
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