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twisted oak moss, twisted screw moss

wideleaf tortula moss

Stems

1–3(–5) mm.

4–12 mm.

Leaves

infolded and somewhat twisted when dry, erect-spreading when moist, spatulate, 1–1.75 × 0.5–0.75 mm;

margins plane (or somewhat erect), crenulate;

apices obtuse, blunt, or emarginate;

costa excurrent into a conspicuously tapered, smooth, yellowish or hyaline awn 1/5–1/2 the length of the leaf, yellow or red, rounded and smooth on abaxial surface;

basal cells rather abruptly differentiated, often with thickened cross-walls, usually extending slightly farther up the margin than near the costa;

distal cells quadrate to hexagonal, 10–15 µm, bulging, with about 4 papillae per cell, the outermost row of cells less papillose.

infolded, twisted, and slightly contorted when dry, erect-spreading when moist, narrowly or broadly spatulate, 1.8–2.8 × 0.6–0.8 mm;

margins revolute in the proximal 1/2, entire;

apices obtuse;

costa ending a few cells before the apex or percurrent, brown, smooth;

basal cells abruptly differentiated, narrower toward the margins;

distal cells rounded, polygonal, or quadrate, 12–18 µm, with 4–6 papillae per cell.

Seta

red, 5–15 mm.

red, 6 mm.

Sexual condition

dioicous or autoicous.

dioicous.

Capsule

red, 3–5 mm, straight, with a distinct neck;

operculum 1–2 mm, red;

peristome 0.5–1 mm, basal membrane 1/2 the total length, red.

red, 3 mm, straight, with a distinct neck;

operculum 1.5–2mm, red;

peristome 1 mm, basal membrane 1/2–2/3 the total length, red.

Spores

10–16 µm, papillose.

10–15 µm, papillose.

Specialized

asexual reproduction as propagula borne at the tip of the stem and in axils of distal leaves, fusiform, green, ecostate, papillose, with an elongate terminal cell, occasionally as several cells.

asexual reproduction by gemmae on the adaxial or occasionally the abaxial leaf surface of leaves, ovoid and rounded, 25–45 µm in diameter, green to brown, multicellular.

Syntrichia laevipila

Syntrichia latifolia

Habitat Occasional on bark of trees, rarely on rock Bark of trees, rarely on rocks
Elevation low to high elevations low to high elevations
Distribution
from FNA
AL; AR; AZ; CA; CO; CT; GA; KS; KY; LA; MD; MO; NC; NE; NM; NV; OH; OK; OR; SC; SD; TN; TX; VA; WA; WV; BC; Mexico; South America (Chile); s Europe; e Asia (Japan); s Africa; Pacific Islands (New Zealand); Australia
[WildflowerSearch map]
from FNA
AZ; CA; OR; WA; BC; Europe
Discussion

This treatment follows M. T. Gallego et al. (2006) in including Syntrichia pagorum with forms of S. laevipila reported in the literature from North America, but this is tentative pending a careful reexamination of this complex of small, corticolous plants. These plants might be confused superfically with S. papillosa, but the leaf margins are plane, the abaxial surface of the costa is smooth and shiny, the propagula are leaf-like, and the leaf cells are pluripapillose in S. laevipila. The other taxa with leaf-like propagula, S. chisosa and S. ammonsiana, are never corticolous.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 27, p. 621. FNA vol. 27, p. 620.
Parent taxa Pottiaceae > subfam. Pottioideae > Syntrichia Pottiaceae > subfam. Pottioideae > Syntrichia
Sibling taxa
S. ammonsiana, S. amphidiacea, S. bartramii, S. cainii, S. caninervis, S. chisosa, S. fragilis, S. latifolia, S. montana, S. norvegica, S. obtusissima, S. papillosa, S. papillosissima, S. princeps, S. ruralis, S. sinensis
S. ammonsiana, S. amphidiacea, S. bartramii, S. cainii, S. caninervis, S. chisosa, S. fragilis, S. laevipila, S. montana, S. norvegica, S. obtusissima, S. papillosa, S. papillosissima, S. princeps, S. ruralis, S. sinensis
Synonyms S. pagorum, Tortula laevipila var. meridionalis, Tortula pagorum Tortula latifolia
Name authority Bridel: Muscol. Recent., suppl. 4: 98. (1818) (Bruch ex Hartman) Huebener: Muscol. Germ., 342. (1833)
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