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Welsh's aster

Drummond's aster

Habit Perennials, 30–100 cm colonial; rhizomes shallow and long, often relatively thick (those of the season each producing a distal rosette near the parent plant). Perennials, 30–120 cm, cespitose; short-rhizomatous or with stout, often woody caudices.
Stems

1–2, ascending to erect (sometimes lax, sometimes reddish), glabrous or sparsely strigoso-villosulous distally (arrays).

1–5+, erect (straight, stout), glabrate or glabrescent, sparsely pilose, or hirsuto-pilose proximally, ± densely so distally.

Leaves

(adaxially green, abaxially pale green, sometimes glaucous) thin (proximal) to stiff (distal), margins serrulate (proximal) or entire or nearly so, ± revolute, scabrous, apices mucronulate or sometimes callous-pointed, faces glabrous;

basal withering by flowering, petiolate (petioles narrowly winged, sheathing), blades oblanceolate to spatulate, 10–40+ × 4–9+ mm, bases attenuate, margins entire, apices obtuse;

proximal cauline withering by flowering, sessile or winged-petiolate, blades lanceolate to oblanceolate, 50–150 × 8–11 mm, bases slightly clasping, margins sparsely serrulate, apices acute;

distal sessile, blades linear to linear-lanceolate, 50–130 × 4–6 mm, gradually reduced distally, bases ± clasping, apices acute.

(abaxially pale or grayish green, adaxially dark green) initially ± firm or membranous, becoming thicker, margins crenate-serrate to serrate, strigoso-ciliate to scabrous, apices acuminate to acute, mucronate, abaxial faces scabrous to densely piloso-strigose, more so on veins, adaxial scabrous to glabrate;

basal withering by flowering, long-petiolate (petioles narrowly winged, sheathing, sometimes ciliate), blades ovate to ovate-oblong or -lanceolate to elliptic or suborbiculate, 10–85(–100) × 8–60 mm, bases deeply cordate to rounded or abruptly attenuate, margins crenate-serrate, apices usually rounded to obtuse, rarely acute;

proximal cauline often withering by flowering, long-petiolate (petioles narrowly winged, more broadly so distally, ciliate), blades ovate or ovate-lanceolate, (25–)50–120(–150) × (15–)20–50(–65) mm, progressively to slightly reduced distally, bases deeply to shallowly cordate or rounded, margins crenate-serrate to sharply serrate;

distal petiolate or subpetiolate or sessile, petioles broadly winged, blades ovate to ovate- or oblong-lanceolate, to oblong-elliptic or linear-lanceolate (4–)5–110 × 1–50 mm, reduced distally (sharply so on array branches in var. texanum), bases shallowly cordate or rounded to attenuate or cuneate (distally), margins crenate-serrate or -serrulate or entire.

Peduncles

0.5–2 cm, scabro-villosulous, bracts linear, scabro-ciliolate.

0, or 0.2–1.5(–4) cm, densely ascending- or appressed-bracteate, pilosulous, bracts 4–17+, lanceolate or subulate to linear-oblanceolate, distally grading into phyllaries, pilosulous or strigoso-ciliate.

Involucres

cylindro-campanulate, 4–7 mm.

campanulate, (3.5–)4.5–6(–6.5) mm.

Ray florets

18–25;

corollas pink to white, laminae 9–12 × 0.8–1.5 mm.

(8–)10–15(—20);

corollas usually bright blue to purple or lavender, sometimes white, laminae (5–)7–10(–12) × 1–1.8 mm.

Disc florets

23–60;

corollas pale yellow, 3.8–6.4 mm, tubes shorter than funnelform throats, lobes narrowly triangular, 0.4–0.8 mm.

(10–)13–15(–22);

corollas cream to yellow, becoming deep purple or reddish purple, 3.5–4.5(–5) mm, tubes shorter than funnelform to cylindric throats, lobes triangular to lanceolate, 0.4–0.7 mm.

Phyllaries

in 3–4 series, lanceolate (outer) to linear (inner), ± unequal, bases indurate, margins hyaline, erose, distally scabro-ciliolate to ciliolate, green zones ± foliaceous (outer) or lanceolate, apices acute to acuminate, sometimes purplish, faces glabrous.

in 4–5 series, subulate or lanceolate (outermost) to narrowly oblong-lanceolate, unequal, bases indurate 1/4–3/4, margins narrowly scarious proximally, hyaline, ciliate, bodies sometimes green to margins distally, green zones lanceolate to linear-lanceolate, apices acute to acuminate or long-acuminate, tips often purple, sometimes involute, mucronulate, faces glabrous.

Heads

in open, slender, leafy, paniculiform or racemiform arrays, branches ascending.

in open, paniculiform arrays, branches ascending to divaricate, often densely racemiform or ± paniculiform (lateral ones open, racemiform) to subsecund, ± stiff to lax, long to relatively short, ± closely to ± remotely leafy, branch leaves progressively or ± abruptly reduced distally.

Cypselae

pinkish tan (nerves stramineous), obovoid, ± compressed, 0.8–1.6 mm, 3–5-nerved, faces sparsely strigillose;

pappi white, 2.8–5.3 mm.

dull purple or brown to tan (ribs tan to stramineous), obovoid, ± compressed, (1.5–)2–3 mm, 5–6-nerved, faces glabrate to sparsely strigillose distally;

pappi cream to rose-tinged, 3–4 mm.

2n

= 16.

Symphyotrichum welshii

Symphyotrichum drummondii

Phenology Flowering Aug–Oct.
Habitat Wet soils in dry areas, hanging gardens, seeps, wet ledges, stream banks
Elevation 1300–2300 m (4300–7500 ft)
Distribution
from FNA
AZ; ID; MT; UT; WY
[BONAP county map]
from FNA
AL; AR; GA; IA; IL; IN; KS; KY; LA; MD; MI; MN; MO; MS; NE; OH; OK; PA; TN; TX; WI; WV; ne Mexico
[WildflowerSearch map]
[BONAP county map]
Discussion

Of conservation concern.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

Variety texanum is tetraploid; var. drummondii is both diploid and tetraploid. The tetraploid race of var. drummondii is most likely autopolyploid; origin of var. texanum has not been determined. The two taxa intergrade and do not appear to warrant species status.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Plants 40–120 cm; stems proximally sparsely hirsute, distally densely so; leaves firm, basal and proximal long-petiolate, petioles ± winged, proximal bases ± cordate or truncate, sometimes obliquely; heads in ample, paniculiform arrays with divaricate or ascending, bracteate, often racemiform branches; peduncles ± secund, 0–4 cm, bracteate; involucres campanulate, (3.5–)4.5–7 mm; ray corollas usually bright blue, light purple, bluish violet, or lavender, sometimes white; cypselae glabrous
var. drummondii
1. Plants 30–80 cm; stems ± densely hirsute, particularly distally; leaves membranous, becoming thickish, brittle, basal and proximal winged-petiolate, proximal bases cordate, becoming rounded-truncate distally; heads in open, paniculiform arrays with very long, widely spreading branches; peduncles 1–4 cm, densely, minutely bracteate; involucres turbinate to hemispheric, 3.8–5.2 mm; ray corollas bluish white; cypselae strigillose
var. texanum
Source FNA vol. 20, p. 523. FNA vol. 20, p. 503.
Parent taxa Asteraceae > tribe Astereae > Symphyotrichum > subg. Symphyotrichum > sect. Symphyotrichum Asteraceae > tribe Astereae > Symphyotrichum > subg. Symphyotrichum > sect. Symphyotrichum
Sibling taxa
S. adnatum, S. anomalum, S. anticostense, S. ascendens, S. boreale, S. bracteolatum, S. campestre, S. chapmanii, S. chilense, S. ciliatum, S. ciliolatum, S. concolor, S. cordifolium, S. cusickii, S. defoliatum, S. depauperatum, S. drummondii, S. dumosum, S. elliottii, S. ericoides, S. eulae, S. falcatum, S. fendleri, S. firmum, S. foliaceum, S. fontinale, S. frondosum, S. georgianum, S. grandiflorum, S. greatae, S. hallii, S. hendersonii, S. jessicae, S. laeve, S. lanceolatum, S. lateriflorum, S. laurentianum, S. lentum, S. molle, S. nahanniense, S. novae-angliae, S. novi-belgii, S. oblongifolium, S. ontarionis, S. oolentangiense, S. parviceps, S. patens, S. phlogifolium, S. pilosum, S. plumosum, S. porteri, S. potosinum, S. praealtum, S. pratense, S. prenanthoides, S. priceae, S. puniceum, S. pygmaeum, S. racemosum, S. retroflexum, S. rhiannon, S. robynsianum, S. sericeum, S. shortii, S. simmondsii, S. spathulatum, S. subspicatum, S. subulatum, S. tenuifolium, S. tradescantii, S. turbinellum, S. undulatum, S. urophyllum, S. walteri, S. yukonense, S. ×amethystinum
S. adnatum, S. anomalum, S. anticostense, S. ascendens, S. boreale, S. bracteolatum, S. campestre, S. chapmanii, S. chilense, S. ciliatum, S. ciliolatum, S. concolor, S. cordifolium, S. cusickii, S. defoliatum, S. depauperatum, S. dumosum, S. elliottii, S. ericoides, S. eulae, S. falcatum, S. fendleri, S. firmum, S. foliaceum, S. fontinale, S. frondosum, S. georgianum, S. grandiflorum, S. greatae, S. hallii, S. hendersonii, S. jessicae, S. laeve, S. lanceolatum, S. lateriflorum, S. laurentianum, S. lentum, S. molle, S. nahanniense, S. novae-angliae, S. novi-belgii, S. oblongifolium, S. ontarionis, S. oolentangiense, S. parviceps, S. patens, S. phlogifolium, S. pilosum, S. plumosum, S. porteri, S. potosinum, S. praealtum, S. pratense, S. prenanthoides, S. priceae, S. puniceum, S. pygmaeum, S. racemosum, S. retroflexum, S. rhiannon, S. robynsianum, S. sericeum, S. shortii, S. simmondsii, S. spathulatum, S. subspicatum, S. subulatum, S. tenuifolium, S. tradescantii, S. turbinellum, S. undulatum, S. urophyllum, S. walteri, S. welshii, S. yukonense, S. ×amethystinum
Subordinate taxa
S. drummondii var. drummondii, S. drummondii var. texanum
Synonyms Aster welshii Aster drummondii, Aster sagittifolius var. drummondii
Name authority (Cronquist) G. L. Nesom: Phytologia 77: 294. (1995) (Lindley) G. L. Nesom: Phytologia 77: 279. (1995)
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