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perennial saltmarsh American-aster, perennial saltmarsh aster

Simmonds' aster

Habit Perennials, (20–)40–60(–100) cm, colonial or cespitose; rhizoma-tous. Perennials, 10–120 cm, colonial (sometimes extensively); long-rhizomatous.
Stems

1–5+, ascending to erect (often with purple or purplish brown areas, flexuous, wiry), little-branched, usually glabrous, sometimes hairy in lines distally.

1–3+, erect (straight, rigid, sometimes stout), sparsely scabrellous or glabrescent to glabrate or glabrous.

Leaves

thick (fleshy), margins entire, faces glabrous;

basal withering by flowering (new winter rosettes appearing at flowering), petiolate (petioles sheathing), blades ovate or oblanceolate, 15–20 × 5–15 mm, bases cuneate, apices rounded;

proximal cauline withering by flowering, usually sessile, blades lanceolate, linear-lanceolate or -oblanceolate, to linear, 7–80(–150) × 1–6 (–12) mm, bases attenuate, margins entire, apices acuminate to acute;

distal sessile, blades narrowly lanceolate to subulate, 10–110 × 0.5–5 mm, apices acuminate.

firm, margins recurved, scabrous to scabrellous, apices acute to acuminate, mucronulate, abaxial faces pilosulous to glabrate or glabrous, midveins marked, adaxial scabridulous;

basal withering by flowering, petiolate (petioles winged, bases sheathing), blades oblanceolate, 26–80 × 6–16 mm, bases attenuate, margins serrate, apices obtuse to acute;

proximal cauline mostly withering by flowering, petiolate to subpetiolate (petioles winged, sheathing to clasping), blades elliptic to oblanceolate or lanceolate or lanceolate-linear to sometimes linear, 45–70+ × 8–15 mm, bases attenuate to cuneate, margins serrate or crenate-serrate or entire;

distal (erect or spreading) sessile, blades lanceolate or oblanceolate to linear, 10–120 × 5–8 mm, reduced distally, sharply so on array branches, bases attenuate to cuneate, margins serrulate or entire.

Peduncles

0.6–4 (–6) cm, bracts 2–8, awl-shaped, grading into phyllaries.

0.3–6+ cm, irregular in length along branches, sometimes distally reduced, glabrate to sparsely scabridulous, bracts 7–12+, spreading to ascending, abruptly reduced distally, grading into phyllaries.

Involucres

narrowly turbinate, 4.1–9.5(–11) mm.

campanulate to campanulo-turbinate, (4.5–)6–8 mm.

Ray florets

10–25;

corollas white or pink, laminae (4.5–)5–8.5(–9.5) × 1.2–2 mm.

18–38;

corollas pale lilac or lavender to pale purple, laminae 9–11 × 0.9–1.7 mm.

Disc florets

25–45(–54);

corollas yellow becoming purplish, 3.4–6(–6.8) mm, tubes shorter than to equaling the narrowly funnelform throats (sparsely hairy at base), lobes ± erect, narrowly triangular, 0.5–0.8 mm.

24–37;

corollas yellow becoming reddish, 4.5–7 mm, tubes shorter than funnelform throats, lobes lanceolate, 0.9–1 mm.

Phyllaries

in 4–5 series, lanceolate to subulate, bases indurate, rounded, margins hyaline, often tinged with purple, entire, green zones spatulate to oblanceolate-rhombic, apices acute, adaxial faces glabrous or minutely hairy distally.

in 5–6 series, oblong-lanceolate to linear-lanceolate or lanceolate (innermost), strongly unequal, bases indurate 1/2–4/5, margins narrowly scarious, distally erose, hyaline, ciliolate, green zones oblanceolate (outer sometimes ± foliaceous), apices acute (inner sometimes acuminate), mucronate, often purplish tipped, faces glabrous.

Heads

(1–)3–20(–40), in open, diffuse, paniculiform arrays, branches patent.

in racemiform to usually open, sometimes ± lax, paniculiform arrays, rarely borne singly, branches ascending to spreading, often straggling and arched, ± leafy (each usually with 10+ heads), secondaries sometimes erect and secund-like.

Cypselae

light brown, narrowly obovoid to fusiform, sometimes ± compressed, 1.5–4(–4.5) mm, 5–6-nerved, faces sparsely strigillose;

pappi tawny to white, 3–6.1 mm.

tan, oblong-obovoid to fusiform-obovoid, compressed, 2–2.6 mm, 4–6-nerved, faces ± strigillose;

pappi cream, 4.6–6.5 mm.

2n

= 64.

Symphyotrichum tenuifolium

Symphyotrichum simmondsii

Phenology Flowering Oct–Jan.
Habitat Moist or dry soils
Elevation 0–50 m (0–200 ft)
Distribution
from FNA
AL; CT; DE; FL; GA; LA; MA; MD; ME; MS; NC; NH; NJ; NY; RI; SC; TX; VA; West Indies
[WildflowerSearch map]
[BONAP county map]
from FNA
FL; NC; SC
[WildflowerSearch map]
Discussion

Varieties 2 (2 in the flora).

Along the Gulf of Mexico Coast in central and northern peninsular Florida, vars. tenuifolium and aphyllum intergrade in nearly all characters (S. D. Sundberg 2004). G. L. Nesom (2005d) treated the varieties as species.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Symphyotrichum simmondsii is known only from peninsular Florida and the southeastern coastal plain. A report from New Jersey is based on an incomplete specimen and appears doubtful.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Stems 1, rhizomes long; mid leaves (1.5–)3–6 mm wide; involucres 6–9.5(–11) mm; ray florets (12–)17–25; disc florets 25–45(–54), corollas (4–)4.7–6(–6.5) mm; cypselae 2.8–4(–4.5) mm; pappi 5–6.1 mm
var. tenuifolium
1. Stems 1–5+ (clustered), rhizomes short; mid leaves (1–)1.5–2.7 mm wide; involucres 4.1–5.3 mm; ray florets 10–16; disc florets (10–)13–23, corollas 3.4–4.6 mm; cypselae 1.5–2(–2.5) mm; pappi 3–4.4 mm
var. aphyllum
Source FNA vol. 20, p. 479. FNA vol. 20, p. 514.
Parent taxa Asteraceae > tribe Astereae > Symphyotrichum > subg. Astropolium Asteraceae > tribe Astereae > Symphyotrichum > subg. Symphyotrichum > sect. Symphyotrichum
Sibling taxa
S. adnatum, S. anomalum, S. anticostense, S. ascendens, S. boreale, S. bracteolatum, S. campestre, S. chapmanii, S. chilense, S. ciliatum, S. ciliolatum, S. concolor, S. cordifolium, S. cusickii, S. defoliatum, S. depauperatum, S. drummondii, S. dumosum, S. elliottii, S. ericoides, S. eulae, S. falcatum, S. fendleri, S. firmum, S. foliaceum, S. fontinale, S. frondosum, S. georgianum, S. grandiflorum, S. greatae, S. hallii, S. hendersonii, S. jessicae, S. laeve, S. lanceolatum, S. lateriflorum, S. laurentianum, S. lentum, S. molle, S. nahanniense, S. novae-angliae, S. novi-belgii, S. oblongifolium, S. ontarionis, S. oolentangiense, S. parviceps, S. patens, S. phlogifolium, S. pilosum, S. plumosum, S. porteri, S. potosinum, S. praealtum, S. pratense, S. prenanthoides, S. priceae, S. puniceum, S. pygmaeum, S. racemosum, S. retroflexum, S. rhiannon, S. robynsianum, S. sericeum, S. shortii, S. simmondsii, S. spathulatum, S. subspicatum, S. subulatum, S. tradescantii, S. turbinellum, S. undulatum, S. urophyllum, S. walteri, S. welshii, S. yukonense, S. ×amethystinum
S. adnatum, S. anomalum, S. anticostense, S. ascendens, S. boreale, S. bracteolatum, S. campestre, S. chapmanii, S. chilense, S. ciliatum, S. ciliolatum, S. concolor, S. cordifolium, S. cusickii, S. defoliatum, S. depauperatum, S. drummondii, S. dumosum, S. elliottii, S. ericoides, S. eulae, S. falcatum, S. fendleri, S. firmum, S. foliaceum, S. fontinale, S. frondosum, S. georgianum, S. grandiflorum, S. greatae, S. hallii, S. hendersonii, S. jessicae, S. laeve, S. lanceolatum, S. lateriflorum, S. laurentianum, S. lentum, S. molle, S. nahanniense, S. novae-angliae, S. novi-belgii, S. oblongifolium, S. ontarionis, S. oolentangiense, S. parviceps, S. patens, S. phlogifolium, S. pilosum, S. plumosum, S. porteri, S. potosinum, S. praealtum, S. pratense, S. prenanthoides, S. priceae, S. puniceum, S. pygmaeum, S. racemosum, S. retroflexum, S. rhiannon, S. robynsianum, S. sericeum, S. shortii, S. spathulatum, S. subspicatum, S. subulatum, S. tenuifolium, S. tradescantii, S. turbinellum, S. undulatum, S. urophyllum, S. walteri, S. welshii, S. yukonense, S. ×amethystinum
Subordinate taxa
S. tenuifolium var. aphyllum, S. tenuifolium var. tenuifolium
Synonyms Aster tenuifolius Aster simmondsii
Name authority (Linnaeus) G. L. Nesom: Phytologia 77: 293. (1995) (Small) G. L. Nesom: Phytologia 77: 291. (1995)
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