Symphyotrichum subulatum |
Symphyotrichum turbinellum |
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annual saltmarsh American-aster, annual saltmarsh aster, annual saltmarsh or eastern annual saltmarsh aster, aster subulé, eastern annual saltmarsh aster, Southwestern annual saltmarsh aster |
prairie aster, smooth violet prairie aster, turbinate aster |
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Habit | Annuals, (10–)30–150 cm; tap-rooted. | Perennials, 30–100 cm, cespitose; with thick, branched, woody caudices. | ||||||||||||||||
Stems | 1, erect (often with purple or purplish brown areas), glabrous or glabrate, sometimes strigillose in leaf axils. |
1–5+, erect, straight (stout, brittle), glabrous or sparsely hirtellous in lines. |
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Leaves | thin (green to dark green), margins often strigilloso-ciliolate, faces glabrous; basal withering by flowering, long-petiolate (petiole bases sheathing), sparsely ciliate, blades ovate to oblanceolate, 10–90 × 6–14 mm, bases attenuate to cuneate, rounded, margins entire or serrulate or crenulate, apices rounded, obtuse, or acute; proximal cauline withering by flowering, petiolate, subpetiolate, or sessile, blades narrowly lanceolate or subulate, 20–100(–200) × 1.5–10(–20) mm, bases attenuate, margins subentire, entire, or serrulate, apices acute to acuminate; distal sessile, blades narrowly lanceolate to subulate, 5–113 × 0.5–5.5 mm, apices acuminate. |
(paler green or bluish green abaxially) firm, margins scabrous, apices mucronate, faces glabrous, abaxial raised midribs scabrous to glabrate; basal withering by flowering, subpetiolate to shortly petiolate (petioles winged, sheathing, coarsely ciliate), blades oblanceolate to oblong-oblanceolate, 45–65 × 5–8 mm, bases cuneate, margins shallowly crenate, strigoso-ciliate, apices acute, obtuse or rounded; proximal cauline withering by flowering, subpetiolate or sessile, blades elliptic-lanceolate to linear-oblanceolate or -lanceolate, 40–120 × 5–20 mm, reduced distally, bases cuneate to slightly attenuate or rounded, slightly clasping, margins serrulate-crenate or entire, apices acute to acuminate; distal sessile, blades oblanceolate or elliptic-lanceolate to linear-lanceolate or linear-oblong, (10–)15–100 × 1–5 mm, gradually reduced distally (more strongly so on branches), bases cuneate or rounded, margins entire, apices acuminate. |
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Peduncles | (0.2–)0.5–4 cm, bracts 4–8(–17). |
long, thin, (1–)4–10(–25) cm, branches to 10 cm, bracts 8–15, regularly spaced but becoming crowded distally, appressed or ascending, linear-oblong to subulate, 1.5–4 mm, grading into phyllaries, mucronulate. |
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Involucres | cylindric to turbinate, 5–7(–8.2) mm. |
turbinate to cylindro-campanulate, 7–12 mm. |
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Ray florets | 16–30(–54) in 1–3 series; corollas white, pink, or lavender, laminae 1.3–7 × 0.2–1.3 mm. |
14–20; corollas light blue to lavender or purple, laminae 12–20 × (1–)1.5–2.8 mm. |
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Disc florets | 4–10(–13); corollas yello, sometimes tinged with purple, 3.4–5.2 mm, throats narrowly funnelform, lobes ± spreading to erect, narrowly triangular, 0.3–0.7 mm, glabrous. |
15–20+; corollas yellow turning purple, (4.5–)5–7 mm, tubes slightly shorter than narrowly funnelform throats, lobes triangular, 0.5–1 mm. |
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Phyllaries | in 3–5 series, broadly or narrowly lanceolate to subulate, unequal, bases indurate, margins hyaline, often purple-tinged, entire, green zones lanceolate (usually narrow, sometimes broad and covering most of distal portion), apices acute, faces glabrous. |
in 6–9 series, appressed or ± spreading, shortly oblong-lanceolate to subulate (outer) to linear-oblong or linear (inner), strongly unequal, bases indurate 1/2–5/6, abaxially rounded, margins scarious, erose, hyaline, sparsely ciliolate distally, green zones oblanceolate to rhombic-oblanceolate, in distal 1/6–1/2, apices acute (outer) to obtuse or rounded (inner), often callous-mucronulate, faces glabrous. |
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Heads | (10–) 30–100(–150), in open, diffuse, paniculiform arrays. |
in open, broad, much ramified, paniculiform arrays, branches ascending to arching, thin, brittle, remotely leafy, rarely sessile. |
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Cypselae | light brown to purple, narrowly obovoid to fusiform, sometimes ± compressed, (1.2–)1.5–2.7(–3) mm, 5-nerved, faces sparsely strigillose; pappi white, (3–)3.5–5.5 mm. |
yellow-tan or light brown to gray, obovoid, ± compressed, (1.8–)2–2.8 mm, 2–4-nerved, faces minutely strigillose; pappi whitish to reddish brown, 4.5–6.4 mm. |
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2n | = 96. |
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Symphyotrichum subulatum |
Symphyotrichum turbinellum |
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Phenology | Flowering Aug–Oct. | |||||||||||||||||
Habitat | Open, dry, acidic (chert, sandstone, or granite), rocky or loamy soils, woods and glades on upland slopes and ridges in soils associated with or on leached soils above bluffs, pastures, roadsides | |||||||||||||||||
Elevation | 60–900 m (200–3000 ft) | |||||||||||||||||
Distribution |
AL; AR; AZ; CA; CT; DE; FL; GA; IL; IN; KS; LA; MA; MD; ME; MI; MS; NC; NE; NH; NJ; NM; NV; NY; OH; OK; PA; RI; SC; TN; TX; UT; VA; NB; ON; Mexico; Central America; South America; West Indies; Bermuda [Widely introduced worldwide]
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AR; IA; IL; KS; LA; MO; NE; OK
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Discussion | Varieties 5 (5 in the flora). Five varieties of Symphyotrichum subulatum are recognized for North America based on differences in chromosome number, ray lamina color and size, array shapes, number of series of ray florets, number of disc and ray florets, and other, more cryptic characters (S. D. Sundberg 2004). These varieties were treated as species by G. L. Nesom (1994b, 2005d). Variety ligulatum is apparently an obligate outcrosser and is the least variable variety (Sundberg). Other varieties are self-compatible, which could facilitate the fixation of mutations in populations. The five varieties are nearly entirely allopatric, and intermediates between pairs of varieties are not uncommon where they approach one another. Populations that are intermediate in ray lamina size between vars. ligulatum and parviflorum are widespread in southern Texas, New Mexico, Arizona, and northern Mexico. Intermediates between vars. elongatum and parviflorum and between vars. elongatum and subulatum occur in Florida. Despite these observations, hybridization experiments and chromosome number differences suggest that the varieties are mostly reproductively isolated (S. D. Sundberg 1986, 2004). In older floras the name Aster exilis Elliott has been applied to Symphyotrichum subulatum vars. ligulatum and parviflorum. The status of this name is uncertain; the type specimen has been lost and the description of the plant is inadequate for determining the taxon to which the name should be applied (G. L. Nesom 1994b; S. D. Sundberg 2004). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Symphyotrichum turbinellum is mostly Ozarkian. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 20, p. 480. | FNA vol. 20, p. 530. | ||||||||||||||||
Parent taxa | Asteraceae > tribe Astereae > Symphyotrichum > subg. Astropolium | Asteraceae > tribe Astereae > Symphyotrichum > subg. Symphyotrichum > sect. Symphyotrichum | ||||||||||||||||
Sibling taxa | ||||||||||||||||||
Subordinate taxa | ||||||||||||||||||
Synonyms | Aster subulatus | Aster turbinellus | ||||||||||||||||
Name authority | (Michaux) G. L. Nesom: Phytologia 77: 293. (1995) | (Lindley) G. L. Nesom: Phytologia 77: 293. (1995) | ||||||||||||||||
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