Symphyotrichum subulatum |
Symphyotrichum racemosum |
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annual saltmarsh American-aster, annual saltmarsh aster, annual saltmarsh or eastern annual saltmarsh aster, aster subulé, eastern annual saltmarsh aster, Southwestern annual saltmarsh aster |
small white American-aster, small white or smooth white Oldfield aster, smooth white Oldfield aster |
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Habit | Annuals, (10–)30–150 cm; tap-rooted. | Perennials, 30–90(–100) cm, colonial or cespitose; long-rhizomatous or with woody caudices. | ||||||||||||||||
Stems | 1, erect (often with purple or purplish brown areas), glabrous or glabrate, sometimes strigillose in leaf axils. |
1–3+, erect (straight), glabrous or glabrate. |
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Leaves | thin (green to dark green), margins often strigilloso-ciliolate, faces glabrous; basal withering by flowering, long-petiolate (petiole bases sheathing), sparsely ciliate, blades ovate to oblanceolate, 10–90 × 6–14 mm, bases attenuate to cuneate, rounded, margins entire or serrulate or crenulate, apices rounded, obtuse, or acute; proximal cauline withering by flowering, petiolate, subpetiolate, or sessile, blades narrowly lanceolate or subulate, 20–100(–200) × 1.5–10(–20) mm, bases attenuate, margins subentire, entire, or serrulate, apices acute to acuminate; distal sessile, blades narrowly lanceolate to subulate, 5–113 × 0.5–5.5 mm, apices acuminate. |
thin, margins often ± revolute, scabrous, apices mucronate to mucronulate, faces glabrous or abaxial minutely pilosulous, cauline with clusters of smaller leaves in most axils; basal withering by flowering (new vernal rosettes often present), petiolate (petioles narrowly winged, sheathing, strigoso-ciliate), blades spatulate to oblanceolate, 5–40 × 5–15 mm, bases attenuate or cuneate to rounded, margins crenate-serrate, apices obtuse to acuminate; proximal cauline withering by flowering, petiolate or subpetiolate (proximalmost) or sessile (petioles winged, sparsely long strigoso-ciliate), blades elliptic or elliptic-lanceolate to lanceolate or linear-lanceolate, 20–70 × 3–20 mm, progressively reduced distally, bases clasping, margins becoming short-ciliate distally; distal (ascending or spreading) usually sessile, sometimes subpetiolate, blades linear-lanceolate to linear, 5–60+ × 1–8 mm, notably unequal in size, reduced distally, abruptly so on branches, bases cuneate to attenuate, margins serrulate or entire. |
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Peduncles | (0.2–)0.5–4 cm, bracts 4–8(–17). |
slender, 0.2–3+ cm or subsessile, hairy in lines, bracts 5–15, linear-elliptic to acicular, 1–2 mm, glabrous, grading into phyllaries. |
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Involucres | cylindric to turbinate, 5–7(–8.2) mm. |
cylindric, (2.5–)3.5–4.5(–5.5) mm. |
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Ray florets | 16–30(–54) in 1–3 series; corollas white, pink, or lavender, laminae 1.3–7 × 0.2–1.3 mm. |
(12–)16–20; corollas usually white, rarely pink, laminae 5–8 × 0.5–1.2 mm. |
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Disc florets | 4–10(–13); corollas yello, sometimes tinged with purple, 3.4–5.2 mm, throats narrowly funnelform, lobes ± spreading to erect, narrowly triangular, 0.3–0.7 mm, glabrous. |
10–20(–25); corollas cream or pale yellow becoming pink or red, (2.5–)3–4.5 mm, tubes shorter than narrowly funnelform throats, lobes recurved to erect, lanceolate, 0.5–1 mm. |
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Phyllaries | in 3–5 series, broadly or narrowly lanceolate to subulate, unequal, bases indurate, margins hyaline, often purple-tinged, entire, green zones lanceolate (usually narrow, sometimes broad and covering most of distal portion), apices acute, faces glabrous. |
in 4–6 series, appressed or outer ± spreading, oblong-lanceolate to linear (innermost) , unequal, bases indurate 1/4–1/2, margins narrowly scarious, hyaline, ciliolate, green zones oblanceolate to linear-oblanceolate, apices acute to acuminate, mucronate, sometimes lightly purple-tinged, faces glabrous. |
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Heads | (10–) 30–100(–150), in open, diffuse, paniculiform arrays. |
in diffuse, ± pyramidal, paniculiform arrays, branches ± lax, spreading horizontally or arching, racemiform, subtended by patent to reflexed branch leaves, often crowded but not (or barely) secund. |
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Cypselae | light brown to purple, narrowly obovoid to fusiform, sometimes ± compressed, (1.2–)1.5–2.7(–3) mm, 5-nerved, faces sparsely strigillose; pappi white, (3–)3.5–5.5 mm. |
gray to tan, obovoid, ± compressed, 1–1.8 mm, 4–5-nerved (faint), faces sparsely strigillose or sericeus; pappi white, 2.5–3.5 mm. |
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2n | = 16. |
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Symphyotrichum subulatum |
Symphyotrichum racemosum |
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Phenology | Flowering Aug–Oct. | |||||||||||||||||
Habitat | Moist to wet, often alluvial soils, often brackish, marshes, savannas, bogs, wet meadows, prairie swales, swamps, borders of swamps, open bottomwoods | |||||||||||||||||
Elevation | 0–200 m (0–700 ft) | |||||||||||||||||
Distribution |
AL; AR; AZ; CA; CT; DE; FL; GA; IL; IN; KS; LA; MA; MD; ME; MI; MS; NC; NE; NH; NJ; NM; NV; NY; OH; OK; PA; RI; SC; TN; TX; UT; VA; NB; ON; Mexico; Central America; South America; West Indies; Bermuda [Widely introduced worldwide]
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AL; AR; CT; DC; DE; FL; GA; IA; IL; IN; KY; LA; MA; MD; ME; MO; NC; NJ; NY; OH; OK; PA; RI; SC; TN; TX; VA; VT; WV; NB; ON
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Discussion | Varieties 5 (5 in the flora). Five varieties of Symphyotrichum subulatum are recognized for North America based on differences in chromosome number, ray lamina color and size, array shapes, number of series of ray florets, number of disc and ray florets, and other, more cryptic characters (S. D. Sundberg 2004). These varieties were treated as species by G. L. Nesom (1994b, 2005d). Variety ligulatum is apparently an obligate outcrosser and is the least variable variety (Sundberg). Other varieties are self-compatible, which could facilitate the fixation of mutations in populations. The five varieties are nearly entirely allopatric, and intermediates between pairs of varieties are not uncommon where they approach one another. Populations that are intermediate in ray lamina size between vars. ligulatum and parviflorum are widespread in southern Texas, New Mexico, Arizona, and northern Mexico. Intermediates between vars. elongatum and parviflorum and between vars. elongatum and subulatum occur in Florida. Despite these observations, hybridization experiments and chromosome number differences suggest that the varieties are mostly reproductively isolated (S. D. Sundberg 1986, 2004). In older floras the name Aster exilis Elliott has been applied to Symphyotrichum subulatum vars. ligulatum and parviflorum. The status of this name is uncertain; the type specimen has been lost and the description of the plant is inadequate for determining the taxon to which the name should be applied (G. L. Nesom 1994b; S. D. Sundberg 2004). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Symphyotrichum racemosum is introduced in Canada. The species is cultivated commercially under the name Aster ericoides cv. ‘Spray’. A. G. Jones (1989) reported hybridization with S. dumosum, S. lateriflorum, S. lanceolatum var. interior, and S. ontarionis. The name Aster vimineus Lamarck has been misapplied to this taxon. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 20, p. 480. | FNA vol. 20, p. 515. | ||||||||||||||||
Parent taxa | Asteraceae > tribe Astereae > Symphyotrichum > subg. Astropolium | Asteraceae > tribe Astereae > Symphyotrichum > subg. Symphyotrichum > sect. Symphyotrichum | ||||||||||||||||
Sibling taxa | ||||||||||||||||||
Subordinate taxa | ||||||||||||||||||
Synonyms | Aster subulatus | Aster racemosus, Aster brachypholis | ||||||||||||||||
Name authority | (Michaux) G. L. Nesom: Phytologia 77: 293. (1995) | (Elliott) G. L. Nesom: Phytologia 77: 290. (1995) | ||||||||||||||||
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