Symphyotrichum subulatum |
Symphyotrichum pygmaeum |
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annual saltmarsh American-aster, annual saltmarsh aster, annual saltmarsh or eastern annual saltmarsh aster, aster subulé, eastern annual saltmarsh aster, Southwestern annual saltmarsh aster |
pygmy aster |
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Habit | Annuals, (10–)30–150 cm; tap-rooted. | Perennials 1.5–15 cm, cespitose; with short, branched caudices, long-rhizomatous (both wiry). | ||||||||||||||||
Stems | 1, erect (often with purple or purplish brown areas), glabrous or glabrate, sometimes strigillose in leaf axils. |
1–10+, decumbent to ascending (purple), sparsely or densely villous to woolly distally. |
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Leaves | thin (green to dark green), margins often strigilloso-ciliolate, faces glabrous; basal withering by flowering, long-petiolate (petiole bases sheathing), sparsely ciliate, blades ovate to oblanceolate, 10–90 × 6–14 mm, bases attenuate to cuneate, rounded, margins entire or serrulate or crenulate, apices rounded, obtuse, or acute; proximal cauline withering by flowering, petiolate, subpetiolate, or sessile, blades narrowly lanceolate or subulate, 20–100(–200) × 1.5–10(–20) mm, bases attenuate, margins subentire, entire, or serrulate, apices acute to acuminate; distal sessile, blades narrowly lanceolate to subulate, 5–113 × 0.5–5.5 mm, apices acuminate. |
firm, margins usually entire, sometimes remotely pauci-serrulate, sparsely villoso-ciliate, apices obtuse to acute, sometimes mucronate; basal often withering by flowering, petiolate (petioles widely winged, sheathing), blades spatulate, 5–19 × 2–4 mm, bases attenuate, apices rounded, faces glabrous or sparsely villous proximally; proximal sessile, blades lanceolate to oblong-lanceolate or (sometimes) -spatulate, 30–50 × 3–10 mm, bases ± clasping, apices obtuse to acute, faces glabrous or sparsely villous; distal sessile, blades lanceolate to oblong, 13–19 × 2–4.5 mm, bases clasping to cuneate, apices acute to obtuse, faces sparsely woolly, sometimes sparsely stipitate-glandular. |
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Peduncles | (0.2–)0.5–4 cm, bracts 4–8(–17). |
densely villous to lanate distally, bracts 0. |
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Involucres | cylindric to turbinate, 5–7(–8.2) mm. |
hemispherico-campanulate, 9–12.5 mm. |
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Ray florets | 16–30(–54) in 1–3 series; corollas white, pink, or lavender, laminae 1.3–7 × 0.2–1.3 mm. |
16–28; corollas purple to violet, laminae 12–18 × 2–3.2 mm. |
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Disc florets | 4–10(–13); corollas yello, sometimes tinged with purple, 3.4–5.2 mm, throats narrowly funnelform, lobes ± spreading to erect, narrowly triangular, 0.3–0.7 mm, glabrous. |
53–55; corollas yellow, 5.6–6.5 mm, throats funnelform, lobes triangular, 0.5–0.8 mm (red or white clavate-hairy). |
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Phyllaries | in 3–5 series, broadly or narrowly lanceolate to subulate, unequal, bases indurate, margins hyaline, often purple-tinged, entire, green zones lanceolate (usually narrow, sometimes broad and covering most of distal portion), apices acute, faces glabrous. |
in 3–4 series (dark purple), lance-oblong or oblong (outer) to linear-lanceolate or sometimes linear (inner), subequal, outer ± herbaceous, bases not indurate, margins herbaceous (outer) to narrowly scarious and erose proximally (inner), strongly purple, villoso-ciliate in green portion, green zones (inner) 1/2–2/3 of distal portions, apices acute to acuminate, inner sometimes apiculate, appressed to loose and squarrose (particularly outer), faces woolly to densely villous, sparsely to moderately stipitate-glandular. |
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Heads | (10–) 30–100(–150), in open, diffuse, paniculiform arrays. |
borne singly. |
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Cypselae | light brown to purple, narrowly obovoid to fusiform, sometimes ± compressed, (1.2–)1.5–2.7(–3) mm, 5-nerved, faces sparsely strigillose; pappi white, (3–)3.5–5.5 mm. |
fusiform to cylindro-obconic, ± compressed, [size unknown], 4–7-nerved (faint), faces ± densely strigillose; pappi whitish to yellowish, 5–7.2 mm. |
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Symphyotrichum subulatum |
Symphyotrichum pygmaeum |
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Phenology | Flowering summer. | |||||||||||||||||
Habitat | Open, active, moist sand dunes, sandy or silty stream banks and terraces, usually cyclically disturbed gravelly tundra and tundra slopes | |||||||||||||||||
Elevation | 0–200+ m (0–700+ ft) | |||||||||||||||||
Distribution |
AL; AR; AZ; CA; CT; DE; FL; GA; IL; IN; KS; LA; MA; MD; ME; MI; MS; NC; NE; NH; NJ; NM; NV; NY; OH; OK; PA; RI; SC; TN; TX; UT; VA; NB; ON; Mexico; Central America; South America; West Indies; Bermuda [Widely introduced worldwide]
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AK; NT; NU |
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Discussion | Varieties 5 (5 in the flora). Five varieties of Symphyotrichum subulatum are recognized for North America based on differences in chromosome number, ray lamina color and size, array shapes, number of series of ray florets, number of disc and ray florets, and other, more cryptic characters (S. D. Sundberg 2004). These varieties were treated as species by G. L. Nesom (1994b, 2005d). Variety ligulatum is apparently an obligate outcrosser and is the least variable variety (Sundberg). Other varieties are self-compatible, which could facilitate the fixation of mutations in populations. The five varieties are nearly entirely allopatric, and intermediates between pairs of varieties are not uncommon where they approach one another. Populations that are intermediate in ray lamina size between vars. ligulatum and parviflorum are widespread in southern Texas, New Mexico, Arizona, and northern Mexico. Intermediates between vars. elongatum and parviflorum and between vars. elongatum and subulatum occur in Florida. Despite these observations, hybridization experiments and chromosome number differences suggest that the varieties are mostly reproductively isolated (S. D. Sundberg 1986, 2004). In older floras the name Aster exilis Elliott has been applied to Symphyotrichum subulatum vars. ligulatum and parviflorum. The status of this name is uncertain; the type specimen has been lost and the description of the plant is inadequate for determining the taxon to which the name should be applied (G. L. Nesom 1994b; S. D. Sundberg 2004). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Symphyotrichum pygmaeum has long been included within or associated with Eurybia sibirica. Though similar in appearance, the two species can be distinguished by the glands present on S. pygmaeum on the distal leaves and phyllaries. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 20, p. 480. | FNA vol. 20, p. 485. | ||||||||||||||||
Parent taxa | Asteraceae > tribe Astereae > Symphyotrichum > subg. Astropolium | Asteraceae > tribe Astereae > Symphyotrichum > subg. Virgulus | ||||||||||||||||
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Subordinate taxa | ||||||||||||||||||
Synonyms | Aster subulatus | Aster pygmaeus, Aster sibiricus subsp. pygmaeus, Aster sibiricus var. pygmaeus, Eurybia pygmaea | ||||||||||||||||
Name authority | (Michaux) G. L. Nesom: Phytologia 77: 293. (1995) | (Lindley) Brouillet & S. Selliah: Sida 21: 1635. (2005) | ||||||||||||||||
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