Symphyotrichum subulatum |
Symphyotrichum phlogifolium |
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annual saltmarsh American-aster, annual saltmarsh aster, annual saltmarsh or eastern annual saltmarsh aster, aster subulé, eastern annual saltmarsh aster, Southwestern annual saltmarsh aster |
thin-leaf late purple aster |
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Habit | Annuals, (10–)30–150 cm; tap-rooted. | Perennials, 50–130 cm, cespitose; with short, thick, woody caudices, not rhizomatous. | ||||||||||||||||
Stems | 1, erect (often with purple or purplish brown areas), glabrous or glabrate, sometimes strigillose in leaf axils. |
1, erect to arching, light to dark brown, soft-hairy, hairs ascending- to spreading- stipitate-glandular distally. |
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Leaves | thin (green to dark green), margins often strigilloso-ciliolate, faces glabrous; basal withering by flowering, long-petiolate (petiole bases sheathing), sparsely ciliate, blades ovate to oblanceolate, 10–90 × 6–14 mm, bases attenuate to cuneate, rounded, margins entire or serrulate or crenulate, apices rounded, obtuse, or acute; proximal cauline withering by flowering, petiolate, subpetiolate, or sessile, blades narrowly lanceolate or subulate, 20–100(–200) × 1.5–10(–20) mm, bases attenuate, margins subentire, entire, or serrulate, apices acute to acuminate; distal sessile, blades narrowly lanceolate to subulate, 5–113 × 0.5–5.5 mm, apices acuminate. |
(dark green) thin, almost membraneous; basal early deciduous, sessile or subpetiolate, blades spatulate to obovate, 80–140 × 20–30 mm, bases cuneate, margins entire to ± serrate, scabrous, apices acute; proximal cauline withering by flowering, sessile, blades panduriform, (20–)30–70(–110) × 10–30(–35) mm, bases auriculate-clasping, margins entire (flat), sparsely villoso-ciliate, apices acute, faces finely sparely villous or scabrous, stipitate-glandular; distal sessile, blades (rugose-veined when fresh) ovate to lanceolate, 75–125(–140) × 15–35(–40) mm, bases strongly cordate-clasping to auriculate-amplexicaul, margins entire (± undulate), apices acute, acuminate to merely mucronate, faces sparsely strigose, moderately so along veins, sparsely to moderately stipitate-glandular. |
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Peduncles | (0.2–)0.5–4 cm, bracts 4–8(–17). |
short (less than 4 cm), strigose, stipitate-glandular, bracts linear-lanceolate, ascending-appressed, grading into phyllaries. |
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Involucres | cylindric to turbinate, 5–7(–8.2) mm. |
cylindric, 5.5–7.5(–8) mm. |
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Ray florets | 16–30(–54) in 1–3 series; corollas white, pink, or lavender, laminae 1.3–7 × 0.2–1.3 mm. |
9–17; corollas light to dark reddish purple, laminae 10–18(–20) × 1–3 mm. |
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Disc florets | 4–10(–13); corollas yello, sometimes tinged with purple, 3.4–5.2 mm, throats narrowly funnelform, lobes ± spreading to erect, narrowly triangular, 0.3–0.7 mm, glabrous. |
15–35; corollas white with purple lobes, 6–8 mm, lobes triangular, 1–1.5 mm. |
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Phyllaries | in 3–5 series, broadly or narrowly lanceolate to subulate, unequal, bases indurate, margins hyaline, often purple-tinged, entire, green zones lanceolate (usually narrow, sometimes broad and covering most of distal portion), apices acute, faces glabrous. |
in 4–6 series, linear-lanceolate, unequal, bases (tan) ± indurate in proximal 1/3–1/2, margins hyaline, erose, distally scabro-ciliolate to ciliolate, green zones covering distal 1/2, apices outer obtuse to acute, inner acuminate, purplish, faces sparsely strigillose, usually moderately to densely stipitate-glandular, rarely sparsely so. |
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Heads | (10–) 30–100(–150), in open, diffuse, paniculiform arrays. |
in crowded, paniculiform arrays, branches often initially patent then ascending. |
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Cypselae | light brown to purple, narrowly obovoid to fusiform, sometimes ± compressed, (1.2–)1.5–2.7(–3) mm, 5-nerved, faces sparsely strigillose; pappi white, (3–)3.5–5.5 mm. |
dark brown to black, obovoid, not compressed, 2.5–4 mm, 7–10-nerved, faces densely strigose along ribs; pappi tan to tawny, sometimes slightly purplish-tinged, 6–8 mm. |
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2n | = 20. |
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Symphyotrichum subulatum |
Symphyotrichum phlogifolium |
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Phenology | Flowering Aug–Oct. | |||||||||||||||||
Habitat | Loamy, sandy, or gravelly soils, rich mesic mixed-hardwood forests (Appalachians), roadsides | |||||||||||||||||
Elevation | 0–1100 m (0–3600 ft) | |||||||||||||||||
Distribution |
AL; AR; AZ; CA; CT; DE; FL; GA; IL; IN; KS; LA; MA; MD; ME; MI; MS; NC; NE; NH; NJ; NM; NV; NY; OH; OK; PA; RI; SC; TN; TX; UT; VA; NB; ON; Mexico; Central America; South America; West Indies; Bermuda [Widely introduced worldwide]
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AL; CT; DC; DE; GA; IN; KY; MA; MD; NC; NY; OH; PA; RI; SC; TN; VA; WV
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Discussion | Varieties 5 (5 in the flora). Five varieties of Symphyotrichum subulatum are recognized for North America based on differences in chromosome number, ray lamina color and size, array shapes, number of series of ray florets, number of disc and ray florets, and other, more cryptic characters (S. D. Sundberg 2004). These varieties were treated as species by G. L. Nesom (1994b, 2005d). Variety ligulatum is apparently an obligate outcrosser and is the least variable variety (Sundberg). Other varieties are self-compatible, which could facilitate the fixation of mutations in populations. The five varieties are nearly entirely allopatric, and intermediates between pairs of varieties are not uncommon where they approach one another. Populations that are intermediate in ray lamina size between vars. ligulatum and parviflorum are widespread in southern Texas, New Mexico, Arizona, and northern Mexico. Intermediates between vars. elongatum and parviflorum and between vars. elongatum and subulatum occur in Florida. Despite these observations, hybridization experiments and chromosome number differences suggest that the varieties are mostly reproductively isolated (S. D. Sundberg 1986, 2004). In older floras the name Aster exilis Elliott has been applied to Symphyotrichum subulatum vars. ligulatum and parviflorum. The status of this name is uncertain; the type specimen has been lost and the description of the plant is inadequate for determining the taxon to which the name should be applied (G. L. Nesom 1994b; S. D. Sundberg 2004). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 20, p. 480. | FNA vol. 20, p. 490. | ||||||||||||||||
Parent taxa | Asteraceae > tribe Astereae > Symphyotrichum > subg. Astropolium | Asteraceae > tribe Astereae > Symphyotrichum > subg. Virgulus | ||||||||||||||||
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Subordinate taxa | ||||||||||||||||||
Synonyms | Aster subulatus | Aster phlogifolius, Aster patens var. phlogifolius, Virgulus patens var. phlogifolius | ||||||||||||||||
Name authority | (Michaux) G. L. Nesom: Phytologia 77: 293. (1995) | (Muhlenberg ex Willdenow) G. L. Nesom: Phytologia 77: 289. (1995) | ||||||||||||||||
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