Symphyotrichum subulatum |
Symphyotrichum oolentangiense |
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annual saltmarsh American-aster, annual saltmarsh aster, annual saltmarsh or eastern annual saltmarsh aster, aster subulé, eastern annual saltmarsh aster, Southwestern annual saltmarsh aster |
azure aster, skyblue aster |
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Habit | Annuals, (10–)30–150 cm; tap-rooted. | Perennials 20–150 cm, cespitose; short-rhizomatous or with branched, woody caudices. | ||||||||||||||||
Stems | 1, erect (often with purple or purplish brown areas), glabrous or glabrate, sometimes strigillose in leaf axils. |
1–5+, erect (straight), glabrate to hispidulous, distally pilosulous to strigillose in arrays. |
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Leaves | thin (green to dark green), margins often strigilloso-ciliolate, faces glabrous; basal withering by flowering, long-petiolate (petiole bases sheathing), sparsely ciliate, blades ovate to oblanceolate, 10–90 × 6–14 mm, bases attenuate to cuneate, rounded, margins entire or serrulate or crenulate, apices rounded, obtuse, or acute; proximal cauline withering by flowering, petiolate, subpetiolate, or sessile, blades narrowly lanceolate or subulate, 20–100(–200) × 1.5–10(–20) mm, bases attenuate, margins subentire, entire, or serrulate, apices acute to acuminate; distal sessile, blades narrowly lanceolate to subulate, 5–113 × 0.5–5.5 mm, apices acuminate. |
thick, firm, margins entire or crenate-serrate, scabrous, apices mucronate, faces scabrous, abaxial strigose to piloso-strigose or hirsute, adaxial strongly strigose; basal withering by flowering, long-petiolate (petioles slightly winged, bases dilated, sheathing, ciliate), blades ovate to lance-ovate, 40–150(–180) × 10–40(–60) mm, bases ± cordate or rounded to attenuate, margins crenate-serrate to subentire, apices rounded or obtuse to acute; proximal cauline petiolate (petioles narrowly winged, clasping), blades ovate or ovate-lanceolate to lanceolate, 30–130 × 12–70 mm, gradually or ± abruptly reduced distally, bases rounded to attenuate, margins entire, scabrous, apices acute or attenuate, callus-pointed; distal sessile or sometimes short-petiolate and ± winged (appressed or ascending), blades lanceolate or linear-lanceolate to subulate, 5–80 × 1–10 mm, strongly reduced distally (more sharply so on branches), bases attenuate to cuneate or ± rounded, margins entire. |
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Peduncles | (0.2–)0.5–4 cm, bracts 4–8(–17). |
ascending, 0.2–6 cm, glabrate, densely bracteate, bracts linear-subulate, scabrous- or ciliate-margined, often revolute, grading with phyllaries. |
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Involucres | cylindric to turbinate, 5–7(–8.2) mm. |
campanulate to cylindro-campanulate, 4.5–8 mm. |
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Ray florets | 16–30(–54) in 1–3 series; corollas white, pink, or lavender, laminae 1.3–7 × 0.2–1.3 mm. |
(10–)13–20(–25); corollas usually pale to azure blue to violet-purple, (rarely rosy, white or bluish white), laminae (5–)8–12(–14) × 1.4–1.7 mm. |
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Disc florets | 4–10(–13); corollas yello, sometimes tinged with purple, 3.4–5.2 mm, throats narrowly funnelform, lobes ± spreading to erect, narrowly triangular, 0.3–0.7 mm, glabrous. |
(15–)20–25(–30); corollas light yellow turning deep purple, 4–5 mm, tubes much shorter than narrowly funnelform throats, lobes triangular to lanceolate, 0.4–0.7 mm (lobes sparsely hairy). |
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Phyllaries | in 3–5 series, broadly or narrowly lanceolate to subulate, unequal, bases indurate, margins hyaline, often purple-tinged, entire, green zones lanceolate (usually narrow, sometimes broad and covering most of distal portion), apices acute, faces glabrous. |
in (3–)4–6 series, appressed or outer slightly spreading, lanceolate to oblong-lanceolate, strongly unequal, bases indurate 1/2–3/4, margins scarious, erose, hyaline, ciliolate, apical green zones diamond-shaped, apices acute to acuminate or caudate (then tips often involute), mucronulate to apiculate, faces glabrous. |
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Heads | (10–) 30–100(–150), in open, diffuse, paniculiform arrays. |
in open, paniculiform arrays, branches ascending, stiff, sometimes long-arching and peduncles secund, ± ridged, densely leafy with small, ± appressed branch leaves, ± strigose. |
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Cypselae | light brown to purple, narrowly obovoid to fusiform, sometimes ± compressed, (1.2–)1.5–2.7(–3) mm, 5-nerved, faces sparsely strigillose; pappi white, (3–)3.5–5.5 mm. |
dull purple or stamineous with purple streaks, oblong-obovoid, ± compressed, 1.8–2 mm, 4–5-nerved, faces glabrate to sparsely strigose; pappi cream or slightly rose-tinged, 3–4 mm. |
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2n | = 32. |
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Symphyotrichum subulatum |
Symphyotrichum oolentangiense |
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Phenology | Flowering Aug–Oct. | |||||||||||||||||
Habitat | Open, dry, sandy, loamy, or rocky soils, dry to wet (seasonally drying) prairies, alvars, glades, bluffs, dunes, barrens, open deciduous woods, oak and/or pine savannas | |||||||||||||||||
Elevation | 50–500 m (200–1600 ft) | |||||||||||||||||
Distribution |
AL; AR; AZ; CA; CT; DE; FL; GA; IL; IN; KS; LA; MA; MD; ME; MI; MS; NC; NE; NH; NJ; NM; NV; NY; OH; OK; PA; RI; SC; TN; TX; UT; VA; NB; ON; Mexico; Central America; South America; West Indies; Bermuda [Widely introduced worldwide]
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AL; AR; FL; GA; IA; IL; IN; KS; LA; MI; MN; MO; MS; NE; NY; OH; OK; PA; SD; TN; TX; WI; ON; Ky ; Mexico (Coahuila)
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Discussion | Varieties 5 (5 in the flora). Five varieties of Symphyotrichum subulatum are recognized for North America based on differences in chromosome number, ray lamina color and size, array shapes, number of series of ray florets, number of disc and ray florets, and other, more cryptic characters (S. D. Sundberg 2004). These varieties were treated as species by G. L. Nesom (1994b, 2005d). Variety ligulatum is apparently an obligate outcrosser and is the least variable variety (Sundberg). Other varieties are self-compatible, which could facilitate the fixation of mutations in populations. The five varieties are nearly entirely allopatric, and intermediates between pairs of varieties are not uncommon where they approach one another. Populations that are intermediate in ray lamina size between vars. ligulatum and parviflorum are widespread in southern Texas, New Mexico, Arizona, and northern Mexico. Intermediates between vars. elongatum and parviflorum and between vars. elongatum and subulatum occur in Florida. Despite these observations, hybridization experiments and chromosome number differences suggest that the varieties are mostly reproductively isolated (S. D. Sundberg 1986, 2004). In older floras the name Aster exilis Elliott has been applied to Symphyotrichum subulatum vars. ligulatum and parviflorum. The status of this name is uncertain; the type specimen has been lost and the description of the plant is inadequate for determining the taxon to which the name should be applied (G. L. Nesom 1994b; S. D. Sundberg 2004). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
A. G. Jones (1989) suggested that Symphyotrichum oolentangiense hybridizes with S. drummondii and S. laeve. Two color morphs, Aster azureus Lindley forma incarnatus Farwell and “forma albidus (the latter not validly published and incorrectly attributed to Steyermark by M. L. Fernald 1950), have been described but are not recognized here. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 20, p. 480. | FNA vol. 20, p. 507. | ||||||||||||||||
Parent taxa | Asteraceae > tribe Astereae > Symphyotrichum > subg. Astropolium | Asteraceae > tribe Astereae > Symphyotrichum > subg. Symphyotrichum > sect. Symphyotrichum | ||||||||||||||||
Sibling taxa | ||||||||||||||||||
Subordinate taxa | ||||||||||||||||||
Synonyms | Aster subulatus | Aster oolentangiensis, Aster azureus, Aster azureus var. poaceus, Aster azureus var. scabrior, Aster capillaceus, Aster poaceus, Aster vernalis | ||||||||||||||||
Name authority | (Michaux) G. L. Nesom: Phytologia 77: 293. (1995) | (Riddell) G. L. Nesom: Phytologia 77: 288. (1995) | ||||||||||||||||
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