Symphyotrichum subulatum |
Symphyotrichum ericoides |
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annual saltmarsh American-aster, annual saltmarsh aster, annual saltmarsh or eastern annual saltmarsh aster, aster subulé, eastern annual saltmarsh aster, Southwestern annual saltmarsh aster |
aster éricoïde, heath American-aster, heath-leaf aster, little gray aster, rayless alkali aster, tuft white prairie aster, white heath aster |
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Habit | Annuals, (10–)30–150 cm; tap-rooted. | Perennials, 20–80(–100) cm, colonial or cespitose, eglandular; branched rhizomatous, or with ± cormoid, branched, woody caudices. | ||||||||||||||||||||
Stems | 1, erect (often with purple or purplish brown areas), glabrous or glabrate, sometimes strigillose in leaf axils. |
1–3+, ascending to erect (grayish brown to brown), sparsely to densely hispido-strigose, sometimes glabrescent proximally. |
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Leaves | thin (green to dark green), margins often strigilloso-ciliolate, faces glabrous; basal withering by flowering, long-petiolate (petiole bases sheathing), sparsely ciliate, blades ovate to oblanceolate, 10–90 × 6–14 mm, bases attenuate to cuneate, rounded, margins entire or serrulate or crenulate, apices rounded, obtuse, or acute; proximal cauline withering by flowering, petiolate, subpetiolate, or sessile, blades narrowly lanceolate or subulate, 20–100(–200) × 1.5–10(–20) mm, bases attenuate, margins subentire, entire, or serrulate, apices acute to acuminate; distal sessile, blades narrowly lanceolate to subulate, 5–113 × 0.5–5.5 mm, apices acuminate. |
usually all except rameal withered by flowering, (light grayish green) firm apices ± white-spine-tipped (often with clusters of smaller leaves in axils); basal sessile, blades (3-nerved) oblanceolate to oblong or spatulate, 10–50 × 10–25 mm, bases attenuate, margins usually entire, rarely remotely serrate, scabrous, apices rounded to obtuse, faces usually sparsely hairy, often glabrous; proximal cauline sessile, blades (1- or 3-nerved) linear to lanceolate or oblong, 10–40(–60) × 1.5–4(–7) mm, reduced distally, bases cuneate, coarsely ciliate, margins entire, coarsely ciliate, apices acute or obtuse, faces moderately to densely strigose or hirsute; distal sessile, blades oblong-ovate, 10–40 × 1.5–3.5 mm, abruptly reduced distally, bases cuneate, margins entire, apices acute, faces moderately to densely strigose. |
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Peduncles | (0.2–)0.5–4 cm, bracts 4–8(–17). |
0.5–1(–2) cm or subsessile, densely hairy, bracts dense, linear to narrowly lanceolate, usually reflexed, sometime appressed to ascending, 1.5–5(–6) mm, densely hairy, grading into phyllaries. |
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Involucres | cylindric to turbinate, 5–7(–8.2) mm. |
cylindric to campanulate, 2.5–4.5(–5) mm. |
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Ray florets | 16–30(–54) in 1–3 series; corollas white, pink, or lavender, laminae 1.3–7 × 0.2–1.3 mm. |
(8–)10–18(–20); corollas usually white, rarely pink or bluish, laminae 6–12(–20) × 0.7–1.2 mm. |
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Disc florets | 4–10(–13); corollas yello, sometimes tinged with purple, 3.4–5.2 mm, throats narrowly funnelform, lobes ± spreading to erect, narrowly triangular, 0.3–0.7 mm, glabrous. |
6–12(–20); corollas yellow becoming brown, 2.5–4 mm, throats narrowly funnelform, lobes triangular, 0.5–0.6 mm, glabrous. |
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Phyllaries | in 3–5 series, broadly or narrowly lanceolate to subulate, unequal, bases indurate, margins hyaline, often purple-tinged, entire, green zones lanceolate (usually narrow, sometimes broad and covering most of distal portion), apices acute, faces glabrous. |
in 3–4 series, oblanceolate to ± spatulate, unequal, firm, bases (whitish to tan) ± indurate in proximal 1/2–2/3, margins hyaline, scabrous proximally, green zones diamond-shaped, in distal 1/2, apices spine-tipped, (outer) spreading to reflexed or squarrose, faces (outer) sparsely to densely hispid, scabroso-hirsute adaxially, (inner) glabrous. |
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Heads | (10–) 30–100(–150), in open, diffuse, paniculiform arrays. |
(1–200+) in paniculiform arrays, branches fastigiate or arrays often pyramidal, racemiform, secund, crowded. |
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Cypselae | light brown to purple, narrowly obovoid to fusiform, sometimes ± compressed, (1.2–)1.5–2.7(–3) mm, 5-nerved, faces sparsely strigillose; pappi white, (3–)3.5–5.5 mm. |
deep purple turning brown, obovoid to oblong-obovoid, ± falcate, not compressed, 1.2–2 × 0.4–0.6 mm, 7–9-nerved (faint), faces sericeous or densely strigillose; pappi whitish, 3–4 mm. |
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Symphyotrichum subulatum |
Symphyotrichum ericoides |
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Distribution |
AL; AR; AZ; CA; CT; DE; FL; GA; IL; IN; KS; LA; MA; MD; ME; MI; MS; NC; NE; NH; NJ; NM; NV; NY; OH; OK; PA; RI; SC; TN; TX; UT; VA; NB; ON; Mexico; Central America; South America; West Indies; Bermuda [Widely introduced worldwide]
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AR; AZ; CO; CT; DC; DE; IA; ID; IL; IN; KS; MA; MD; ME; MI; MN; MO; MS; MT; ND; NE; NJ; NM; NY; OH; OK; OR; PA; RI; SD; TX; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NT; ON; QC; SK; n Mexico
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Discussion | Varieties 5 (5 in the flora). Five varieties of Symphyotrichum subulatum are recognized for North America based on differences in chromosome number, ray lamina color and size, array shapes, number of series of ray florets, number of disc and ray florets, and other, more cryptic characters (S. D. Sundberg 2004). These varieties were treated as species by G. L. Nesom (1994b, 2005d). Variety ligulatum is apparently an obligate outcrosser and is the least variable variety (Sundberg). Other varieties are self-compatible, which could facilitate the fixation of mutations in populations. The five varieties are nearly entirely allopatric, and intermediates between pairs of varieties are not uncommon where they approach one another. Populations that are intermediate in ray lamina size between vars. ligulatum and parviflorum are widespread in southern Texas, New Mexico, Arizona, and northern Mexico. Intermediates between vars. elongatum and parviflorum and between vars. elongatum and subulatum occur in Florida. Despite these observations, hybridization experiments and chromosome number differences suggest that the varieties are mostly reproductively isolated (S. D. Sundberg 1986, 2004). In older floras the name Aster exilis Elliott has been applied to Symphyotrichum subulatum vars. ligulatum and parviflorum. The status of this name is uncertain; the type specimen has been lost and the description of the plant is inadequate for determining the taxon to which the name should be applied (G. L. Nesom 1994b; S. D. Sundberg 2004). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 2 (2 in the flora). Symphyotrichum ericoides resembles S. pilosum var. pilosum, which has larger heads, longer rays, and phyllaries that are not spine-tipped, though the revolute margins can make them appear so. Two subspecies and four weakly separated varieties of S. ericoides were recognized by A. G. Jones (1978). Tetraploids of var. ericoides on the eastern prairies can be difficult to distinguish from S. falcatum. A number of aster cultivars are sold under the name “Aster ericoides.” These are all derived from European garden plants and are either cultivars of S. dumosum, S. lateriflorum, S. pilosum, or S. racemosum, or hybrids involving one of those species and another taxon. The misapplication of the epithet ericoides dates back to the nineteenth century and has persisted in the horticultural literature. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 20, p. 480. | FNA vol. 20, p. 494. | ||||||||||||||||||||
Parent taxa | Asteraceae > tribe Astereae > Symphyotrichum > subg. Astropolium | Asteraceae > tribe Astereae > Symphyotrichum > subg. Virgulus | ||||||||||||||||||||
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Synonyms | Aster subulatus | Aster ericoides, Lasallea ericoides, Virgulus ericoides | ||||||||||||||||||||
Name authority | (Michaux) G. L. Nesom: Phytologia 77: 293. (1995) | (Linnaeus) G. L. Nesom: Phytologia 77: 280. (1995) | ||||||||||||||||||||
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