Symphyotrichum subulatum |
Symphyotrichum elliottii |
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annual saltmarsh American-aster, annual saltmarsh aster, annual saltmarsh or eastern annual saltmarsh aster, aster subulé, eastern annual saltmarsh aster, Southwestern annual saltmarsh aster |
Elliott's aster, marsh American-aster |
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Habit | Annuals, (10–)30–150 cm; tap-rooted. | Perennials, 60–160(–200) cm, colonial; long-rhizomatous. | ||||||||||||||||
Stems | 1, erect (often with purple or purplish brown areas), glabrous or glabrate, sometimes strigillose in leaf axils. |
1, erect (straight, stout, sometimes purplish pink), mostly glabrous, hirsutulous in arrays, mainly in lines. |
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Leaves | thin (green to dark green), margins often strigilloso-ciliolate, faces glabrous; basal withering by flowering, long-petiolate (petiole bases sheathing), sparsely ciliate, blades ovate to oblanceolate, 10–90 × 6–14 mm, bases attenuate to cuneate, rounded, margins entire or serrulate or crenulate, apices rounded, obtuse, or acute; proximal cauline withering by flowering, petiolate, subpetiolate, or sessile, blades narrowly lanceolate or subulate, 20–100(–200) × 1.5–10(–20) mm, bases attenuate, margins subentire, entire, or serrulate, apices acute to acuminate; distal sessile, blades narrowly lanceolate to subulate, 5–113 × 0.5–5.5 mm, apices acuminate. |
firm, margins serrate to serrulate, scabridulous, apices acute to shortly acuminate, apiculate to mucronate, abaxial faces glabrous, adaxial scabridulous; basal withering by flowering, long-petiolate (petioles narrowly winged, sheathing), blades elliptic, 50–250 × 10–50 mm, bases attenuate; proximal cauline withering by flowering, long-petiolate to subpetiolate (petioles winged, bases expanded, sheathing), blades elliptic to lanceolate or oblanceolate, gradually reduced distally, 70–110 × 15–35 mm, bases attenuate; distal subpetiolate or sessile, (petioles broadly winged, bases sheathing to clasping), blades elliptic, 10–70 × 1–30 mm, progressively reduced distally, more strongly so on branches, bases attenuate to cuneate, clasping, margins serrulate or entire. |
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Peduncles | (0.2–)0.5–4 cm, bracts 4–8(–17). |
0.5–2.5 cm, to erect, hirsute, bracts 3–5, linear, grading into phyllaries. |
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Involucres | cylindric to turbinate, 5–7(–8.2) mm. |
narrowly campanulate, 8–11 mm. |
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Ray florets | 16–30(–54) in 1–3 series; corollas white, pink, or lavender, laminae 1.3–7 × 0.2–1.3 mm. |
(25–)30–46+; corollas pink or sometimes lavender, laminae 7–14 × 0.8–1.6 mm. |
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Disc florets | 4–10(–13); corollas yello, sometimes tinged with purple, 3.4–5.2 mm, throats narrowly funnelform, lobes ± spreading to erect, narrowly triangular, 0.3–0.7 mm, glabrous. |
25–35+; corollas pale yellow becoming pink and later brown, 6–6.2 mm, tubes slightly shorter than narrowly funnelform limbs, lobes narrowly triangular to lanceolate, 0.8–1 mm. |
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Phyllaries | in 3–5 series, broadly or narrowly lanceolate to subulate, unequal, bases indurate, margins hyaline, often purple-tinged, entire, green zones lanceolate (usually narrow, sometimes broad and covering most of distal portion), apices acute, faces glabrous. |
in 5–6 series, linear-lanceolate or -oblanceolate to linear, slightly unequal, bases indurate 1/3–1/2, margins narrowly scarious, hyaline, sparsely and remotely ciliolate, green zones lanceolate to linear, outer distally foliaceous, sometimes constricted in middle, apices spreading to squarrose, long-acuminate, apiculate, faces glabrous. |
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Heads | (10–) 30–100(–150), in open, diffuse, paniculiform arrays. |
in paniculiform arrays, branches strongly ascending, leafy. |
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Cypselae | light brown to purple, narrowly obovoid to fusiform, sometimes ± compressed, (1.2–)1.5–2.7(–3) mm, 5-nerved, faces sparsely strigillose; pappi white, (3–)3.5–5.5 mm. |
tan to pale brown, oblanceoloid, compressed, 2–2.7 mm, 3–4-nerved, faces glabrous or sparsely hairy; pappi sordid or whitish, 5.5–5.8 mm. |
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2n | = 16. |
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Symphyotrichum subulatum |
Symphyotrichum elliottii |
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Phenology | Flowering Aug–Oct. | |||||||||||||||||
Habitat | Moist, low sites, swamps, bogs, marshes, brackish marshes | |||||||||||||||||
Elevation | 0–50+ m (0–200+ ft) | |||||||||||||||||
Distribution |
AL; AR; AZ; CA; CT; DE; FL; GA; IL; IN; KS; LA; MA; MD; ME; MI; MS; NC; NE; NH; NJ; NM; NV; NY; OH; OK; PA; RI; SC; TN; TX; UT; VA; NB; ON; Mexico; Central America; South America; West Indies; Bermuda [Widely introduced worldwide]
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AL; FL; GA; LA; NC; SC; VA
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Discussion | Varieties 5 (5 in the flora). Five varieties of Symphyotrichum subulatum are recognized for North America based on differences in chromosome number, ray lamina color and size, array shapes, number of series of ray florets, number of disc and ray florets, and other, more cryptic characters (S. D. Sundberg 2004). These varieties were treated as species by G. L. Nesom (1994b, 2005d). Variety ligulatum is apparently an obligate outcrosser and is the least variable variety (Sundberg). Other varieties are self-compatible, which could facilitate the fixation of mutations in populations. The five varieties are nearly entirely allopatric, and intermediates between pairs of varieties are not uncommon where they approach one another. Populations that are intermediate in ray lamina size between vars. ligulatum and parviflorum are widespread in southern Texas, New Mexico, Arizona, and northern Mexico. Intermediates between vars. elongatum and parviflorum and between vars. elongatum and subulatum occur in Florida. Despite these observations, hybridization experiments and chromosome number differences suggest that the varieties are mostly reproductively isolated (S. D. Sundberg 1986, 2004). In older floras the name Aster exilis Elliott has been applied to Symphyotrichum subulatum vars. ligulatum and parviflorum. The status of this name is uncertain; the type specimen has been lost and the description of the plant is inadequate for determining the taxon to which the name should be applied (G. L. Nesom 1994b; S. D. Sundberg 2004). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Symphyotrichum elliottii grows on the Atlantic coastal plain. It is of conservation concern in some states. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 20, p. 480. | FNA vol. 20, p. 524. | ||||||||||||||||
Parent taxa | Asteraceae > tribe Astereae > Symphyotrichum > subg. Astropolium | Asteraceae > tribe Astereae > Symphyotrichum > subg. Symphyotrichum > sect. Symphyotrichum | ||||||||||||||||
Sibling taxa | ||||||||||||||||||
Subordinate taxa | ||||||||||||||||||
Synonyms | Aster subulatus | Aster elliottii | ||||||||||||||||
Name authority | (Michaux) G. L. Nesom: Phytologia 77: 293. (1995) | (Torrey & A. Gray) G. L. Nesom: Phytologia 77: 280. (1995) | ||||||||||||||||
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