Symphyotrichum subulatum |
Symphyotrichum defoliatum |
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annual saltmarsh American-aster, annual saltmarsh aster, annual saltmarsh or eastern annual saltmarsh aster, aster subulé, eastern annual saltmarsh aster, Southwestern annual saltmarsh aster |
San Bernardino aster |
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Habit | Annuals, (10–)30–150 cm; tap-rooted. | Perennials, (30–)40–100(–150) cm, cespitose; short-rhizomatous. | ||||||||||||||||
Stems | 1, erect (often with purple or purplish brown areas), glabrous or glabrate, sometimes strigillose in leaf axils. |
1–5+, ascending to erect, strigose, villous, or lanate, especially distally. |
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Leaves | thin (green to dark green), margins often strigilloso-ciliolate, faces glabrous; basal withering by flowering, long-petiolate (petiole bases sheathing), sparsely ciliate, blades ovate to oblanceolate, 10–90 × 6–14 mm, bases attenuate to cuneate, rounded, margins entire or serrulate or crenulate, apices rounded, obtuse, or acute; proximal cauline withering by flowering, petiolate, subpetiolate, or sessile, blades narrowly lanceolate or subulate, 20–100(–200) × 1.5–10(–20) mm, bases attenuate, margins subentire, entire, or serrulate, apices acute to acuminate; distal sessile, blades narrowly lanceolate to subulate, 5–113 × 0.5–5.5 mm, apices acuminate. |
(grayish green) thin to firm, margins entire, apices acute, faces strigose; basal withering by flowering, petiolate, blades narrowly oblanceolate, 50–100(–150) × 4–12(–15) mm, bases attenuate, margins scabrous to ciliate; proximalmost cauline often withering by flowering (often with tufts of leaves in axils), sessile, blades linear to narrowly oblong, 30–70(–100) × 4–8(–10) mm, bases attenuate or cuneate, apices acute; distal sessile, blades 15–80 × 2–8(–10) mm, bases cuneate, sometimes auriculate. |
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Peduncles | (0.2–)0.5–4 cm, bracts 4–8(–17). |
densely short-strigose, bracts 1–6, linear-oblong, strigose. |
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Involucres | cylindric to turbinate, 5–7(–8.2) mm. |
campanulate, 4–7 mm. |
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Ray florets | 16–30(–54) in 1–3 series; corollas white, pink, or lavender, laminae 1.3–7 × 0.2–1.3 mm. |
15–40; corollas pale violet, laminae 8–12(–15) × 1–1.5(–2) mm. |
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Disc florets | 4–10(–13); corollas yello, sometimes tinged with purple, 3.4–5.2 mm, throats narrowly funnelform, lobes ± spreading to erect, narrowly triangular, 0.3–0.7 mm, glabrous. |
25–70+; corollas yellow, 4–6 mm, lobes triangular, 0.4–0.8 mm. |
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Phyllaries | in 3–5 series, broadly or narrowly lanceolate to subulate, unequal, bases indurate, margins hyaline, often purple-tinged, entire, green zones lanceolate (usually narrow, sometimes broad and covering most of distal portion), apices acute, faces glabrous. |
in 3–6 series, narrowly oblanceolate or linear (outer) to linear (inner), unequal, bases indurate, margins entire, narrowly scarious, ciliate, green zones obovate to elliptic, apices obtuse, mucronulate, faces usually strigose. |
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Heads | (10–) 30–100(–150), in open, diffuse, paniculiform arrays. |
(3–50+) in narrowly paniculiform arrays, branches 2–10(–20) cm. |
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Cypselae | light brown to purple, narrowly obovoid to fusiform, sometimes ± compressed, (1.2–)1.5–2.7(–3) mm, 5-nerved, faces sparsely strigillose; pappi white, (3–)3.5–5.5 mm. |
brown, cylindric to obovoid, not compressed, 1.5–3 mm, ribs 5–8, faces hairy; pappi whitish, 4–6 mm. |
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2n | = 36. |
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Symphyotrichum subulatum |
Symphyotrichum defoliatum |
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Phenology | Flowering Aug–Nov. | |||||||||||||||||
Habitat | Dry open grasslands and meadows, often near springs | |||||||||||||||||
Elevation | 0–1700 m (0–5600 ft) | |||||||||||||||||
Distribution |
AL; AR; AZ; CA; CT; DE; FL; GA; IL; IN; KS; LA; MA; MD; ME; MI; MS; NC; NE; NH; NJ; NM; NV; NY; OH; OK; PA; RI; SC; TN; TX; UT; VA; NB; ON; Mexico; Central America; South America; West Indies; Bermuda [Widely introduced worldwide]
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CA
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Discussion | Varieties 5 (5 in the flora). Five varieties of Symphyotrichum subulatum are recognized for North America based on differences in chromosome number, ray lamina color and size, array shapes, number of series of ray florets, number of disc and ray florets, and other, more cryptic characters (S. D. Sundberg 2004). These varieties were treated as species by G. L. Nesom (1994b, 2005d). Variety ligulatum is apparently an obligate outcrosser and is the least variable variety (Sundberg). Other varieties are self-compatible, which could facilitate the fixation of mutations in populations. The five varieties are nearly entirely allopatric, and intermediates between pairs of varieties are not uncommon where they approach one another. Populations that are intermediate in ray lamina size between vars. ligulatum and parviflorum are widespread in southern Texas, New Mexico, Arizona, and northern Mexico. Intermediates between vars. elongatum and parviflorum and between vars. elongatum and subulatum occur in Florida. Despite these observations, hybridization experiments and chromosome number differences suggest that the varieties are mostly reproductively isolated (S. D. Sundberg 1986, 2004). In older floras the name Aster exilis Elliott has been applied to Symphyotrichum subulatum vars. ligulatum and parviflorum. The status of this name is uncertain; the type specimen has been lost and the description of the plant is inadequate for determining the taxon to which the name should be applied (G. L. Nesom 1994b; S. D. Sundberg 2004). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Symphyotrichum defoliatum is known from the San Gabriel and San Bernardino mountains, and from the Peninsular Ranges, southern California. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 20, p. 480. | FNA vol. 20, p. 538. | ||||||||||||||||
Parent taxa | Asteraceae > tribe Astereae > Symphyotrichum > subg. Astropolium | Asteraceae > tribe Astereae > Symphyotrichum > subg. Ascendentes | ||||||||||||||||
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Synonyms | Aster subulatus | Aster defoliatus, Aster bernardinus, Aster chilensis var. bernardinus | ||||||||||||||||
Name authority | (Michaux) G. L. Nesom: Phytologia 77: 293. (1995) | (Parish) G. L. Nesom: Phytologia 77: 279. (1995) | ||||||||||||||||
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