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enable glossary links

annual saltmarsh American-aster, annual saltmarsh aster, annual saltmarsh or eastern annual saltmarsh aster, aster subulé, eastern annual saltmarsh aster, Southwestern annual saltmarsh aster

aster boréal, northern bog aster, rush American-aster, rush aster, rush or slender white or northern bog aster, slender white aster

Habit Annuals, (10–)30–150 cm; tap-rooted. Perennials (5.5–)13–85 cm, colonial; slender (0.6–2.8 mm diam.), long-rhizomatous (rhizomes of season shallow- or deep-seated, not producing rosettes near parent stems).
Stems

1, erect (often with purple or purplish brown areas), glabrous or glabrate, sometimes strigillose in leaf axils.

1–3+, erect (straight, slender, often reddish to slightly glaucous proximally), glabrous.

Leaves

thin (green to dark green), margins often strigilloso-ciliolate, faces glabrous;

basal withering by flowering, long-petiolate (petiole bases sheathing), sparsely ciliate, blades ovate to oblanceolate, 10–90 × 6–14 mm, bases attenuate to cuneate, rounded, margins entire or serrulate or crenulate, apices rounded, obtuse, or acute;

proximal cauline withering by flowering, petiolate, subpetiolate, or sessile, blades narrowly lanceolate or subulate, 20–100(–200) × 1.5–10(–20) mm, bases attenuate, margins subentire, entire, or serrulate, apices acute to acuminate;

distal sessile, blades narrowly lanceolate to subulate, 5–113 × 0.5–5.5 mm, apices acuminate.

stiff, margins entire to sparsely subserrulate, revolute, scabrellous, apices mucronulate, sometimes with callous point, faces glabrous, abaxial midveins sometimes sparsely strigillose;

basal withering by flowering, subpetiolate to petiolate, petioles winged, base dilated, sheathing, eciliate, blades linear-oblanceolate to ovate or suborbiculate, 20–150 × 5–60 mm, bases attenuate, margins shallowly serrate or entire, apices obtuse to rounded;

proximal cauline withering by flowering, sessile or subpetiolate, petioles widely winged, bases strongly clasping, blades linear-lanceolate to linear, 15–90 × 2–12 mm, bases attenuate to cuneate or subauriculate, ± clasping, apices obtuse to acute;

distal sessile, blades linear-lanceolate (sometimes lanceolate) to linear, (40–)60–150 × 2–6(–8) mm, ± reduced distally, bases subauriculate, subclasping, margins entire or sometimes remotely serrulate, apices acute.

Peduncles

(0.2–)0.5–4 cm, bracts 4–8(–17).

0.5–5 cm, glabrous, bracts 1–3, remote, linear-lanceolate to linear, reduced distally, not grading into phyllaries (sometimes subtending heads).

Involucres

cylindric to turbinate, 5–7(–8.2) mm.

cylindro-campanulate, (4–)5–8 mm.

Ray florets

16–30(–54) in 1–3 series;

corollas white, pink, or lavender, laminae 1.3–7 × 0.2–1.3 mm.

(15–)25–35(–41);

corollas white to often pale rose, pale purple or lavender, laminae (7–)10–15(–20) × 1.1–1.5 mm.

Disc florets

4–10(–13);

corollas yello, sometimes tinged with purple, 3.4–5.2 mm, throats narrowly funnelform, lobes ± spreading to erect, narrowly triangular, 0.3–0.7 mm, glabrous.

(15–)25–30(–40);

corollas cream or pale yellow becoming pink or brown purple, 3.7–6.6 mm, tubes shorter than funnelform throats, lobes triangular, 0.5–1 mm.

Phyllaries

in 3–5 series, broadly or narrowly lanceolate to subulate, unequal, bases indurate, margins hyaline, often purple-tinged, entire, green zones lanceolate (usually narrow, sometimes broad and covering most of distal portion), apices acute, faces glabrous.

in 4–5 series, usually appressed, rarely outer recurved, lanceolate or oblong-lanceolate (outer) to linear (innermost) ± unequal to sometimes subequal, bases indurate 1/3–1/2 (sometimes not indurate), margins scarious, erose, hyaline or tinged with reddish purple, sparsely ciliolate, green zones lanceolate to linear-lanceolate, apices acute to acuminate, mucronate to apiculate, often purple or reddish purple (particularly inner), faces glabrous.

Heads

(10–) 30–100(–150), in open, diffuse, paniculiform arrays.

borne singly or in open, often lax, racemiform or paniculiform arrays, branches ascending, sparsely leafy.

Cypselae

light brown to purple, narrowly obovoid to fusiform, sometimes ± compressed, (1.2–)1.5–2.7(–3) mm, 5-nerved, faces sparsely strigillose;

pappi white, (3–)3.5–5.5 mm.

yellowish tan or purple to brownish with purple streaks or grayish tan (nerves stramineous), obovoid, ± compressed, 0.6–2 mm, 3–5-nerved, faces sparsely strigillose;

pappi white, 2.7–6.4 mm.

2n

= 16, 32, 48, 64.

Symphyotrichum subulatum

Symphyotrichum boreale

Phenology Flowering Aug–Oct.
Habitat Mostly calcareous areas, fens, marshes, bogs, open cedar-tamarack-spruce swamps, stream and pond margins, wet meadows, swales
Elevation 0–1500+ m (0–4900+ ft)
Distribution
from FNA
AL; AR; AZ; CA; CT; DE; FL; GA; IL; IN; KS; LA; MA; MD; ME; MI; MS; NC; NE; NH; NJ; NM; NV; NY; OH; OK; PA; RI; SC; TN; TX; UT; VA; NB; ON; Mexico; Central America; South America; West Indies; Bermuda [Widely introduced worldwide]
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; CO; IA; ID; IL; IN; MA; MI; MN; MT; ND; NE; NH; NJ; NY; OH; PA; SD; VT; WA; WI; WV; WY; AB; BC; MB; NB; NF; NS; NT; NU; ON; PE; QC; SK; YT
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 5 (5 in the flora).

Five varieties of Symphyotrichum subulatum are recognized for North America based on differences in chromosome number, ray lamina color and size, array shapes, number of series of ray florets, number of disc and ray florets, and other, more cryptic characters (S. D. Sundberg 2004). These varieties were treated as species by G. L. Nesom (1994b, 2005d). Variety ligulatum is apparently an obligate outcrosser and is the least variable variety (Sundberg). Other varieties are self-compatible, which could facilitate the fixation of mutations in populations.

The five varieties are nearly entirely allopatric, and intermediates between pairs of varieties are not uncommon where they approach one another. Populations that are intermediate in ray lamina size between vars. ligulatum and parviflorum are widespread in southern Texas, New Mexico, Arizona, and northern Mexico. Intermediates between vars. elongatum and parviflorum and between vars. elongatum and subulatum occur in Florida. Despite these observations, hybridization experiments and chromosome number differences suggest that the varieties are mostly reproductively isolated (S. D. Sundberg 1986, 2004).

In older floras the name Aster exilis Elliott has been applied to Symphyotrichum subulatum vars. ligulatum and parviflorum. The status of this name is uncertain; the type specimen has been lost and the description of the plant is inadequate for determining the taxon to which the name should be applied (G. L. Nesom 1994b; S. D. Sundberg 2004).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Some of the western populations originally determined as Symphyotrichum boreale in western Wyoming and adjacent Montana are referable to S. welshii. Those of eastern Wyoming, Colorado, and the Black Hills are S. boreale. There is a gap between the ranges of the two species in Wyoming where neither occurs. Symphyotrichum ×longulum (E. Sheldon) G. L. Nesom (syn. Aster longulus E. Sheldon), reported from Minnesota, is the hybrid between S. boreale and S. puniceum var. puniceum.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Ray laminae lavender to blue, (3.5–)4.5–7 × 0.9–1.3 mm, drying in 3–5 coils; ray florets in 1 series; disc florets (20–)33–50 (sc United States)
var. ligulatum
1. Ray laminae white to pink or lavender, 1.3–3.5(–4.2) × 0.2–0.6 mm, drying in (0–)1–3(–4) coils; ray florets in 1–3 series; disc florets 3–23
→ 2
2. Ray laminae pink to lavender, (2–)2.5–3.5(–4.2) × 0.3–0.6 mm, drying in 2–3(–4) coils; disc florets 11–23 (Florida, West Indies)
var. elongatum
2. Ray laminae usually white, rarely pink or lavender, 1.3–3 × 0.2–0.5 mm, drying in 1 coil or curling; disc florets (3–)4–15
→ 3
3. Ray laminae longer than pappi; phyllaries 30–42; pappi 3.5–3.8(–4.2) mm; w United States
var. parviflorum
3. Ray laminae shorter to slightly longer than pappi; phyllaries 18–30; pappi 3.5–5.5 mm; e and se United States
→ 4
4. Green zones of phyllaries narrowly to broadly lanceolate, extending phyllary length; ray laminae 0.2–0.5 mm wide; disc florets 3.8–4.6(–4.9) mm; salt marshes, e North America and Gulf Coast
var. subulatum
4. Green zones of phyllaries broadly lanceolate, nearly absent from proximal portion; ray laminae 0.2–0.3 mm wide; disc florets 3.2–4.1 mm; adventive, s United States
var. squamatum
Source FNA vol. 20, p. 480. FNA vol. 20, p. 522.
Parent taxa Asteraceae > tribe Astereae > Symphyotrichum > subg. Astropolium Asteraceae > tribe Astereae > Symphyotrichum > subg. Symphyotrichum > sect. Symphyotrichum
Sibling taxa
S. adnatum, S. anomalum, S. anticostense, S. ascendens, S. boreale, S. bracteolatum, S. campestre, S. chapmanii, S. chilense, S. ciliatum, S. ciliolatum, S. concolor, S. cordifolium, S. cusickii, S. defoliatum, S. depauperatum, S. drummondii, S. dumosum, S. elliottii, S. ericoides, S. eulae, S. falcatum, S. fendleri, S. firmum, S. foliaceum, S. fontinale, S. frondosum, S. georgianum, S. grandiflorum, S. greatae, S. hallii, S. hendersonii, S. jessicae, S. laeve, S. lanceolatum, S. lateriflorum, S. laurentianum, S. lentum, S. molle, S. nahanniense, S. novae-angliae, S. novi-belgii, S. oblongifolium, S. ontarionis, S. oolentangiense, S. parviceps, S. patens, S. phlogifolium, S. pilosum, S. plumosum, S. porteri, S. potosinum, S. praealtum, S. pratense, S. prenanthoides, S. priceae, S. puniceum, S. pygmaeum, S. racemosum, S. retroflexum, S. rhiannon, S. robynsianum, S. sericeum, S. shortii, S. simmondsii, S. spathulatum, S. subspicatum, S. tenuifolium, S. tradescantii, S. turbinellum, S. undulatum, S. urophyllum, S. walteri, S. welshii, S. yukonense, S. ×amethystinum
S. adnatum, S. anomalum, S. anticostense, S. ascendens, S. bracteolatum, S. campestre, S. chapmanii, S. chilense, S. ciliatum, S. ciliolatum, S. concolor, S. cordifolium, S. cusickii, S. defoliatum, S. depauperatum, S. drummondii, S. dumosum, S. elliottii, S. ericoides, S. eulae, S. falcatum, S. fendleri, S. firmum, S. foliaceum, S. fontinale, S. frondosum, S. georgianum, S. grandiflorum, S. greatae, S. hallii, S. hendersonii, S. jessicae, S. laeve, S. lanceolatum, S. lateriflorum, S. laurentianum, S. lentum, S. molle, S. nahanniense, S. novae-angliae, S. novi-belgii, S. oblongifolium, S. ontarionis, S. oolentangiense, S. parviceps, S. patens, S. phlogifolium, S. pilosum, S. plumosum, S. porteri, S. potosinum, S. praealtum, S. pratense, S. prenanthoides, S. priceae, S. puniceum, S. pygmaeum, S. racemosum, S. retroflexum, S. rhiannon, S. robynsianum, S. sericeum, S. shortii, S. simmondsii, S. spathulatum, S. subspicatum, S. subulatum, S. tenuifolium, S. tradescantii, S. turbinellum, S. undulatum, S. urophyllum, S. walteri, S. welshii, S. yukonense, S. ×amethystinum
Subordinate taxa
S. subulatum var. elongatum, S. subulatum var. ligulatum, S. subulatum var. parviflorum, S. subulatum var. squamatum, S. subulatum var. subulatum
Synonyms Aster subulatus Aster laxifolius var. borealis, Aster borealis, Aster franklinianus, Aster junciformis
Name authority (Michaux) G. L. Nesom: Phytologia 77: 293. (1995) (Torrey & A. Gray) Á. Löve & D. Löve: Taxon 31: 358. (1982)
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