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rigid whitetop aster

alkali American aster, aster cilié, rayless alkali aster, rayless annual American-aster, rayless annual aster, rayless annual or rayless alkali aster

Habit Perennials, 40–100 cm, usually colonial, sometimes ± cespitose; short-(seldom long-)rhizomatous. Annuals, 7–70+ cm.
Stems

1–2+, erect (straight to slightly flexuous, sometimes robust), glabrous.

1, ascending to erect (straight), bluish yellowish green, often red-tinged, ± succulent, glabrous.

Leaves

thin to firm, margins slightly recurved, scabrous, faces glabrous;

basal mostly withering by flowering, petiolate (petioles narrowly winged, sheathing), blades lanceolate-ovate to ovate or spatulate, 25–110 × 10–40 mm, bases usually rounded to attenuate, seldom ± cordate, margins serrulate to shallowly serrate, apices obtuse to acute;

proximal cauline often withering by flowering, petiolate (petioles ± widely winged, ± clasping), blades lance-ovate or narrowly ovate, 50–95 × 19–31 mm, bases usually rounded to attenuate, sometimes cuneate, rarely subcordate, margins serrate to serrulate;

distal winged-petiolate or subpetiolate or sessile, blades lance-ovate or narrowly ovate to lanceolate or sometimes linear (in arrays), 7–1.5 × 3–31 mm, gradually and strongly reduced distally, bases cuneate, scarcely or not clasping, to slightly rounded and ± clasping, margins serrulate or entire, apices acute to acuminate.

bluish (green) thin, sometimes ± fleshy, margins usually entire, sometimes serrulate, strigoso-ciliate to scabrous, midribs conspicuous, apices acute to short-acuminate, faces glabrous;

basal withering by flowering, petiolate, blades spatulate, 15–205 × 1.5–9 mm, bases attenuate;

proximal cauline usually withering by flowering (with clusters of smaller leaves in axils, often elongating into branches);

cauline sessile, blades linear-oblanceolate, (10–)30–80(–150) × 1–4(–9) mm, gradually reduced distally, bases slightly dilated and clasping to rounded.

Peduncles

(when present) 0.2–1.5 cm, densely pilose or pilosulous, bracts 1–6+, lanceolate to oblong-lanceolate, foliaceous to distally grading into phyllaries.

Involucres

campanulate, (6–)7–10 mm.

narrowly campanulate, 5–7(–11) mm.

Ray florets

13–30;

corollas blue to purple, laminae (10–)15–17 × 1.6–3 mm.

0.

Pistillate florets

75–95+ in 4–5+ series;

laminae 0 (corolla tubes ± 2 mm, shorter than style branches).

Disc florets

35–50+;

corollas cream to pale yellow becoming pinkish, 5.8–6.8 mm, tubes shorter than funnelform throats, lobes lanceolate, 0.9–1.2 mm, glabrous.

± 14;

corollas whitish turning pink, ± ampliate, tubes slender, longer than narrowly funnelform limbs, 3.5–5 mm, lobes narrowly triangular, ± 0.2 mm.

Phyllaries

in 3–4(–5) series, broadly lanceolate constricted to narrow oblong bases (outer) to oblong-lanceolate or linear-lanceolate (inner), ± unequal, bases indurate 1/10–1/5 (outer) to 4/5+ (inner), margins hyaline, narrowly scarious, erose (indurate portion), sparsely ciliolate (foliaceous portion), green zones foliaceous (outer) to distal 1/5–1/4 foliaceous (inner), basally truncate to lanceolate, apices spreading to squarrose, acute to acuminate, mucronulate, abaxial faces glabrous or glabrate, adaxial strigillose (foliaceous portion).

in 3–4 series, loose, linear to narrowly oblanceolate, subequal or outer sometimes longer, bases scarious, margins narrowly scarious proximally (outer), scabrous, green zones foliaceous (outer and mid) to lanceolate (inner), apices acute (rarely obtuse), mucronulate, faces glabrous.

Heads

in broad (appearing ± corymbiform) or narrow, paniculiform to ± racemiform arrays, branches usually long (to 10 cm), ascending to divaricate.

(disciform) in ± dense, narrow to pyramidal, paniculiform to racemiform arrays, branches decumbent (proximal) to ascending (distal);

peduncles 0–1 cm, glabrous, bracts linear-lanceolate to linear, crowding heads.

Cypselae

tan, obovoid to oblanceoloid, compressed, 3–3.9 mm, 4–6-nerved, faces glabrous or sparsely strigillose;

pappi whitish to reddish, 5.3–6 mm.

purple or grayish with purple streaks, obovoid to oblong-obovoid, ± compressed, 1.5–2.5 mm, 2–4-nerved (faint), faces hirsuto-strigose;

pappi white or pinkish, 4–6 mm.

2n

= 48.

= 14.

Symphyotrichum retroflexum

Symphyotrichum ciliatum

Phenology Flowering Aug–Oct. Flowering Aug–Oct.
Habitat Moist meadows, moist woodlands, dry, open, pine or oak woods, wooded roadsides Moist, brackish soils, prairies, steppes, salt marshes (Hudson Bay), summer-receding prairie ponds, open grounds in loess hills, irrigation channels, winter-salted highways, railroads, waste grounds
Elevation 400–1500 m (1300–4900 ft) 0–2000+ m (0–6600+ ft)
Distribution
from FNA
AL; GA; NC; SC; TN; VA
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; CO; IA; ID; IL; IN; KS; ME; MI; MN; MO; MT; ND; NE; NY; OH; OK; PA; SD; UT; WA; WI; WY; AB; BC; MB; NB; NT; ON; QC; SK; YT; n Eurasia (w to Rumania)
[WildflowerSearch map]
[BONAP county map]
Discussion

Symphyotrichum retroflexum is known only from the Appalachian Blue Ridges. It is possible that the species is an allopolyploid derived from members of subsections Cordifolii and Symphyotrichum.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Symphyotrichum ciliatum is introduced east of the Prairies in winter-salted wastegrounds and roadsides; it is native in northern Ontario, however, in the saltmarshes of the western shore of James Bay. It is native to the steppes of Eurasia, westward to Rumania.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 20, p. 510. FNA vol. 20, p. 499.
Parent taxa Asteraceae > tribe Astereae > Symphyotrichum > subg. Symphyotrichum > sect. Symphyotrichum Asteraceae > tribe Astereae > Symphyotrichum > subg. Symphyotrichum > sect. Conyzopsis
Sibling taxa
S. adnatum, S. anomalum, S. anticostense, S. ascendens, S. boreale, S. bracteolatum, S. campestre, S. chapmanii, S. chilense, S. ciliatum, S. ciliolatum, S. concolor, S. cordifolium, S. cusickii, S. defoliatum, S. depauperatum, S. drummondii, S. dumosum, S. elliottii, S. ericoides, S. eulae, S. falcatum, S. fendleri, S. firmum, S. foliaceum, S. fontinale, S. frondosum, S. georgianum, S. grandiflorum, S. greatae, S. hallii, S. hendersonii, S. jessicae, S. laeve, S. lanceolatum, S. lateriflorum, S. laurentianum, S. lentum, S. molle, S. nahanniense, S. novae-angliae, S. novi-belgii, S. oblongifolium, S. ontarionis, S. oolentangiense, S. parviceps, S. patens, S. phlogifolium, S. pilosum, S. plumosum, S. porteri, S. potosinum, S. praealtum, S. pratense, S. prenanthoides, S. priceae, S. puniceum, S. pygmaeum, S. racemosum, S. rhiannon, S. robynsianum, S. sericeum, S. shortii, S. simmondsii, S. spathulatum, S. subspicatum, S. subulatum, S. tenuifolium, S. tradescantii, S. turbinellum, S. undulatum, S. urophyllum, S. walteri, S. welshii, S. yukonense, S. ×amethystinum
S. adnatum, S. anomalum, S. anticostense, S. ascendens, S. boreale, S. bracteolatum, S. campestre, S. chapmanii, S. chilense, S. ciliolatum, S. concolor, S. cordifolium, S. cusickii, S. defoliatum, S. depauperatum, S. drummondii, S. dumosum, S. elliottii, S. ericoides, S. eulae, S. falcatum, S. fendleri, S. firmum, S. foliaceum, S. fontinale, S. frondosum, S. georgianum, S. grandiflorum, S. greatae, S. hallii, S. hendersonii, S. jessicae, S. laeve, S. lanceolatum, S. lateriflorum, S. laurentianum, S. lentum, S. molle, S. nahanniense, S. novae-angliae, S. novi-belgii, S. oblongifolium, S. ontarionis, S. oolentangiense, S. parviceps, S. patens, S. phlogifolium, S. pilosum, S. plumosum, S. porteri, S. potosinum, S. praealtum, S. pratense, S. prenanthoides, S. priceae, S. puniceum, S. pygmaeum, S. racemosum, S. retroflexum, S. rhiannon, S. robynsianum, S. sericeum, S. shortii, S. simmondsii, S. spathulatum, S. subspicatum, S. subulatum, S. tenuifolium, S. tradescantii, S. turbinellum, S. undulatum, S. urophyllum, S. walteri, S. welshii, S. yukonense, S. ×amethystinum
Synonyms Aster retroflexus, Aster curtisii Erigeron ciliatus, Aster angustus, Aster brachyactis, Brachyactis angusta, Brachyactis ciliata subsp. angusta
Name authority (Lindley ex de Candolle) G. L. Nesom: Phytologia 77: 291. (1995) (Ledebour) G. L. Nesom: Phytologia 77: 277. (1995)
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