Symphyotrichum puniceum var. scabricaule |
Symphyotrichum puniceum |
|||||
---|---|---|---|---|---|---|
purplestem aster, roughstem aster |
aster ponceau, purple-stem American-aster, purple-stem aster, purplestem or swamp or red-stem aster, swamp aster |
|||||
Habit | Plants 100–250(–300) cm. | Perennials (7–)100–250(–300) cm, cespitose; thickly short-rhizomatous or with stout, branched caudices, sometimes long-rhizomatous. | ||||
Stems | usually densely and uniformly hirsute, sometimes less so and in lines distally. |
1–5+, erect (straight to distally flexuous, stout 5–11 mm diam., usually reddish purple), usually densely to sparsely hirsute or hispid, glabrescent, often proximally glabrate, sometimes glabrate. |
||||
Leaves | faces ± concolorous, without dark, distinct reticulum, adaxial with distinctly impressed main veins (giving rough appearance); array leaves reduced in size relative to mid cauline. |
(dark green, not crowded) thin, margins remotely serrate to sometimes entire, scabrous, apices acute or acuminate to subcaudate, mucronate, abaxial faces glabrate, ± densely pilose along midveins, adaxial glabrate to scabrous; basal withering by flowering, subpetiolate to petiolate (petioles ± winged, bases dilated, sheathing), blades spatulate to oblanceolate or lanceolate, 30–10+ × 3–20+ mm, bases attenuate to cuneate, margins remotely crenate-serrate to denticulate or entire, apices acute or acuminate to rounded; proximal cauline withering by flowering, petiolate to subpetiolate (petioles widely winged, bases auriculate, clasping), blades lanceolate or oblanceolate to oblong, elliptic or elliptic-oblong, (55–)70–200(–220) × 10–40 mm, usually progressively lengthened to mid, bases ± attenuate to cuneate or auriculate and clasping; distal sessile, blades oblanceolate or lanceolate to oblong or elliptic-oblong (sometimes nearly panduriform), 8–160 × 1–40 mm, progressively reduced distally, strongly so and linear in arrays, bases ± strongly auriculate, clasping, margins subentire or entire. |
||||
Peduncles | 0.2–3+ cm, ± hispid or hirsute to villous, bracts 0–3, lanceolate or linear-lanceolate to linear, distal often subtending heads. |
|||||
Involucres | campanulate, (6–)8–12(–15) mm. |
|||||
Ray florets | 20–50(–60); corollas usually pale to dark blue, lavender or purple, seldom white or pink, laminae (7–)12–18(–21) × (0.9–)1.4–1.8 mm. |
|||||
Disc florets | 30–50 (–90); corollas yellow or cream becoming pink or purple, (4.1–)5–6.5 mm, tubes shorter than cylindro-funnelform limbs, lobes triangular, 0.4–1.1 mm. |
|||||
Phyllaries | in 4–6 series, oblanceolate-spatulate to linear, subequal, bases indurate 1/5–1/2, margins not scarious (outer) to narrowly scarious, erose, hyaline, sparsely ciliolate, green zones linear-lanceolate, outer sometimes ± foliaceous, to linear (inner), apices ± spreading at tip when foliaceous, long-acuminate to caudate, mucronate to apiculate, sometimes purplish tinged, faces glabrous or ± hispid. |
|||||
Heads | in open, paniculiform (often corymbiform-looking) arrays, branches widely spreading to ascending, sparsely to moderately leafy with ± large leaves. |
|||||
Cypselae | purple or brown (nerves stramineous), oblanceoloid, ± falcate, compressed, (2–)2.5–3.5(–4) mm, 3–4-nerved (nerves prominent), faces glabrous or sparsely strigillose; pappi white, 3.8–6 mm. |
|||||
2n | = 16. |
|||||
Symphyotrichum puniceum var. scabricaule |
Symphyotrichum puniceum |
|||||
Phenology | Flowering Sep–Oct(–Nov). | |||||
Habitat | Open seepage sites with mucky, saturated soils, wet, seepy grounds in sandy pinelands, sphagnum bogs, shrubby seepage bogs, marshes, pond margins, open stream banks, roadside and drainage ditches | |||||
Elevation | 0–100+ m (0–300+ ft) | |||||
Distribution |
AL; LA; MS; TX |
AL; CT; DE; GA; IA; IL; IN; LA; MD; ME; MI; MN; MO; MS; NC; ND; NE; NH; NJ; NY; OH; PA; RI; SC; SD; TN; TX; VA; VT; WI; WV; AB; BC; MB; NB; NS; ON; PE; QC; SK
|
||||
Discussion | Of conservation concern. Variety scabricaule is known from the post-oak belt of eastern Texas and in Louisiana (Natchitoches Parish), Mississippi (Grenada and Lauderdale counties), and Alabama (Penton, Chambers County, Kral 37887, BRIT). It is considered of conservation concern in Texas, where most populations occur. Variety scabricaule is in the Center for Plant Conservation’s National Collection of Endangered Plants. The hybrid Symphyotrichum puniceum var. scabricaule × S. lateriflorum was reported by G. L. Nesom (1997b). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 2 (2 in the flora). Symphyotrichum puniceum is one of the widest ranging species in the genus, growing from the edges of the prairies to the Atlantic coast, and from the Gulf coast of Texas to southern Ungava Bay (northern Quebec). Several taxa have been recognized within this variable species. Here, we segregate Symphyotrichum firmum as a distinct entity. The tall, pink-rayed S. elliottii of the outer coastal plains is also distinctive, despite reports of hybrids with S. puniceum var. puniceum where their ranges overlap (see G. L. Nesom 1997b). A recently described member of the complex, S. rhiannon, is an ultramafic endemic from North Carolina. Here, within S. puniceum, we recognize two varieties, the variable and widespread var. puniceum, and var. scabricaule from the inner Gulf coastal plains, following Nesom, who also offered a useful summary of the infraspecific taxonomy of the species. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
||||
Key |
|
|||||
Source | FNA vol. 20, p. 525. | FNA vol. 20, p. 524. | ||||
Parent taxa | ||||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Aster scabricaulis, Aster puniceus var. scabricaulis | Aster puniceus | ||||
Name authority | (Shinners) G. L. Nesom: Phytologia 77: 290. (1995) | (Linnaeus) Á. Löve & D. Löve: Taxon 31: 359. (1982) | ||||
Web links |
|