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willow aster, willow-leaf American-aster, willowleaf aster

aster poilu, awl American-aster, frost aster, hairy aster, hairy white Oldfield aster, Oldfield or frost weed or white Oldfield or hairy or hairy white Oldfield aster, white heath aster

Habit Perennials (10–)50–150(–200) cm, colonial; fleshy long-rhizomatous. Perennials cespitose, (5–)20–120(–150+) cm; with stout, branched caudices, sometimes long-rhizomatous.
Stems

1, erect to ascending (stout, ± glaucous or reddish), glabrate to moderately hispidulous-hirsute (mostly in lines) distally.

1–5+, ascending to erect (straight, stout), glabrate to hirsuto-hispid or pilose, sometimes glabrous or hairy in lines (var. pringlei).

Leaves

thick, firm, margins often revolute, scabrous, apices mucronate, abaxial faces glabrous, vein areoles conspicuous, ± isodiametric, adaxial waxy, glabrate to strigose, ± scabrous (distal cauline with axillary leaf clusters, sometimes becoming branches);

basal withering by flowering, petiolate to subpetiolate (petioles winged, bases sheathing, ciliate), blades spatulate, 40–70 × 10–25 mm, bases attenuate, margins entire to shallowly serrate, apices rounded to obtuse;

proximal cauline withering by flowering, sessile (subpetiolate), blades elliptic or lanceolate to oblanceolate or linear-lanceolate, 40–100(–150) × 3–18 mm, progressively reduced distally, bases cuneate, often slightly rounded, apices acute to attenuate, callus-pointed, abaxial faces scabrellous, adaxial glabrous or ± puberulent;

distal sessile, blades ovate or elliptic-lanceolate to lanceolate or linear, 10–85 × 2–17 mm, progressively reduced distally, bases cuneate, margins entire, apices acute or obtuse, callus-pointed.

thin, margins ciliate, apices acute to acuminate, mucronate, faces glabrous or ± hirsute (particularly on abaxial midveins);

basal withering by flowering (new vernal rosettes developing at flowering), petiolate to subpetiolate (petioles winged, ciliate, bases sheathing), blades oblanceolate or obovate to spatulate, 10–60 × 5–15 mm, bases attenuate, margins sparsely crenate-serrate (mostly apically), apices obtuse to rounded;

proximal cauline usually deciduous at flowering (often with axillary clusters of small leaves), petiolate or subpetiolate to subsessile (petioles narrowly to broadly winged, clasping), blades elliptic-oblanceolate or -oblong to linear-lanceolate or -oblanceolate, 40–102 × 5–25 mm, progressively reduced distally, bases attenuate to cuneate, ± clasping, margins entire to serrate, softly ciliate, apices attenuate, hyaline-spinulose;

distal sessile, blades lance-oblong to linear-lanceolate to linear or linear-oblanceolate or linear-subulate, 10–100 × 1–8 mm, progressively reduced distally, branch leaves abruptly smaller, bases cuneate, margins entire or serrulate.

Peduncles

0.3–2 cm, ± pilose, bracts 5–12+, ascending or recurved-spreading, lance-elliptic to lanceolate or linear-lanceolate, foliaceous, distal closely subtending heads, not grading into phyllaries.

5–30(–50) mm, progressively shorter distally, glabrous or densely hispid, bracts 7–25+, appressed to ascending, sometimes ± spreading, linear to subulate, glabrate, ciliate or not, mucronate to apiculate, ± grading into phyllaries.

Involucres

campanulate, (4–)5–7(–8) mm.

campanulate to cylindro-campanulate, (2.5–)3.5–5.1(–6.5) mm.

Ray florets

(6–)20–35;

corollas pale blue-violet to lavender or rose-purple, rarely white, laminae 5–10(–12) × 1–1.7 mm.

(10–)16–28(–38);

corollas usually white, rarely pinkish or bluish, laminae (4–)5.4–7.5(–11) × (0.4–)0.8–1.3(–1.7) mm.

Disc florets

20–30(–35+);

corollas cream or light yellow turning pinkish purple, 4–6.5 mm, tubes shorter than narrowly funnelform throats, lobes lanceolate, 0.5–1.3 mm.

(13–)17–39(–67);

corollas light yellow becoming reddish purple or brown, (2.5–)3–4.1(–5.5) mm, tubes much shorter than narrowly funnelform throats, lobes lanceolate, (0.4–)0.6–0.8(–1) mm.

Phyllaries

in 4–6 series, appressed, oblong-lanceolate to linear (innermost) (outer sometimes ± foliaceous), unequal, bases indurate 1/2–3/4, margins scarious, erose, hyaline, sparsely ciliolate (often ciliate basally), green zones lanceolate to lance-rhombic, apices sometimes spreading, acute to acuminate, mucronate, often reddish purple apically, faces glabrous.

in 4–6 series, appressed or slightly spreading, oblong-lanceolate (outer) to linear (innermost) , unequal (rarely subequal), bases indurate 1/2–3/5, margins hyaline, scarious, erose, ± ciliolate distally, green zones lanceolate to lance-rhombic, apices acute or acuminate, involute, spinulose, faces glabrous or sparsely hirsutulous.

Heads

in ample, foliaceous, usually dense, paniculiform arrays, branches ascending to spreading, ± racemiform.

in open, leafy, often pyramidal, paniculiform arrays, branches divaricate or branches 10 cm or less ascending, often arched and secund, sometimes racemiform.

Cypselae

purple or stramineous with purple, obovoid, sometimes ± compressed, 1.5–2 mm, 4–5-nerved, faces thinly strigillose;

pappi white, 4–6.5 mm.

whitish or gray, oblong-obovoid, sometimes ± compressed, 1–1.5 × 0.5–0.7 mm, 4–6-nerved, faces sparsely strigillose;

pappi white, 3.5–4 mm.

2n

= 32, 48, 64.

Symphyotrichum praealtum

Symphyotrichum pilosum

Phenology Flowering Aug–Oct.
Habitat Wet, loamy soils, wet prairies or meadows, lake and stream shores, oak savannas, open woods or thickets, fields, moist banks, ditches, roadsides, recent clearings
Elevation 0–400+ m (0–1300+ ft)
Distribution
from FNA
AL; AR; CT; DC; DE; FL; IA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MO; MS; NE; NH; NJ; NY; OH; OK; PA; RI; SD; TN; TX; VA; WI; WV; ON; Mexico (Chihuahua, Coahuila, Nuevo Léon) [Introduced in Europe]
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; CT; DC; DE; FL; GA; IA; IL; IN; KS; KY; MA; MD; ME; MI; MN; MO; MS; NC; NE; NH; NJ; NY; OH; OK; PA; RI; SC; SD; TN; VA; VT; WI; WV; BC; NB; NS; ON; QC
[WildflowerSearch map]
[BONAP county map]
Discussion

Symphyotrichum praealtum was reported by H. R. Hinds (2000) as a possible introduced, non-persisting ephemeral in New Brunswick. The species is introduced in central Europe, where it has been confused with S. lanceolatum (J. C. Semple et al. 2002). A. G. Jones (1989) reported hybridization with S. lanceolatum and S. firmum.

G. L. Nesom (1994b, 1997) and J. C. Semple et al. (2002) recognize several varieties within the species: var. praealtum [syn. Aster coerulescens de Candolle, A. praealtus var. coerulescens (de Candolle) A. G. Jones, A. praealtus var. imbricatior Wiegand] (2n = 32); var. angustior (Wiegand) G. L. Nesom (syn. A. praealtus var. angustior Wiegand) (2n = 64); var. nebraskense (Britton) G. L. Nesom (syn. A. praealtus var. nebraskensis Britton); var. subasperum (Lindley) G. L. Nesom [syn. A. praealtus var. subasper (Lindley) Wiegand, A. subasper Lindley]; and var. texicola (Wiegand) G. L. Nesom (syn. A. praealtus var. texicola Wiegand). Some of these varieties deserve recognition, but the validity of others needs confirmation.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

The ecology and cytogeography of Symphyotrichum pilosum were summarized by J. G. Chmielewski and J. C. Semple (2001b). The species is often confused with S. ericoides, which inhabits the same kinds of open, prairie-like habitats; the latter can be distinguished by its smaller heads and spine-tipped phyllary apices. Because of that, the name Aster ericoides has been misapplied often in floras and in the horticultural trade.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Stems sparsely to densely hirsute; leaves pilose; various substrates
var. pilosum
1. Stems glabrous; leaves glabrous or glabrate; mainly on calcareous substrates
var. pringlei
Source FNA vol. 20, p. 521. FNA vol. 20, p. 511.
Parent taxa Asteraceae > tribe Astereae > Symphyotrichum > subg. Symphyotrichum > sect. Symphyotrichum Asteraceae > tribe Astereae > Symphyotrichum > subg. Symphyotrichum > sect. Symphyotrichum
Sibling taxa
S. adnatum, S. anomalum, S. anticostense, S. ascendens, S. boreale, S. bracteolatum, S. campestre, S. chapmanii, S. chilense, S. ciliatum, S. ciliolatum, S. concolor, S. cordifolium, S. cusickii, S. defoliatum, S. depauperatum, S. drummondii, S. dumosum, S. elliottii, S. ericoides, S. eulae, S. falcatum, S. fendleri, S. firmum, S. foliaceum, S. fontinale, S. frondosum, S. georgianum, S. grandiflorum, S. greatae, S. hallii, S. hendersonii, S. jessicae, S. laeve, S. lanceolatum, S. lateriflorum, S. laurentianum, S. lentum, S. molle, S. nahanniense, S. novae-angliae, S. novi-belgii, S. oblongifolium, S. ontarionis, S. oolentangiense, S. parviceps, S. patens, S. phlogifolium, S. pilosum, S. plumosum, S. porteri, S. potosinum, S. pratense, S. prenanthoides, S. priceae, S. puniceum, S. pygmaeum, S. racemosum, S. retroflexum, S. rhiannon, S. robynsianum, S. sericeum, S. shortii, S. simmondsii, S. spathulatum, S. subspicatum, S. subulatum, S. tenuifolium, S. tradescantii, S. turbinellum, S. undulatum, S. urophyllum, S. walteri, S. welshii, S. yukonense, S. ×amethystinum
S. adnatum, S. anomalum, S. anticostense, S. ascendens, S. boreale, S. bracteolatum, S. campestre, S. chapmanii, S. chilense, S. ciliatum, S. ciliolatum, S. concolor, S. cordifolium, S. cusickii, S. defoliatum, S. depauperatum, S. drummondii, S. dumosum, S. elliottii, S. ericoides, S. eulae, S. falcatum, S. fendleri, S. firmum, S. foliaceum, S. fontinale, S. frondosum, S. georgianum, S. grandiflorum, S. greatae, S. hallii, S. hendersonii, S. jessicae, S. laeve, S. lanceolatum, S. lateriflorum, S. laurentianum, S. lentum, S. molle, S. nahanniense, S. novae-angliae, S. novi-belgii, S. oblongifolium, S. ontarionis, S. oolentangiense, S. parviceps, S. patens, S. phlogifolium, S. plumosum, S. porteri, S. potosinum, S. praealtum, S. pratense, S. prenanthoides, S. priceae, S. puniceum, S. pygmaeum, S. racemosum, S. retroflexum, S. rhiannon, S. robynsianum, S. sericeum, S. shortii, S. simmondsii, S. spathulatum, S. subspicatum, S. subulatum, S. tenuifolium, S. tradescantii, S. turbinellum, S. undulatum, S. urophyllum, S. walteri, S. welshii, S. yukonense, S. ×amethystinum
Subordinate taxa
S. pilosum var. pilosum, S. pilosum var. pringlei
Synonyms Aster praealtus, Aster salicifolius Aster pilosus, Aster ericoides var. pilosus
Name authority (Poiret) G. L. Nesom: Phytologia 77: 289. (1995) (Willdenow) G. L. Nesom: Phytologia 77: 289. (1995)
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