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Santa Rita Mountain aster

aster cordifolié, common blue wood aster, heart-leaf American-aster, heart-leaf aster, heartleaf or common blue wood aster

Habit Perennials, 15–35(–45) cm, colonial or cespitose; rhizoma-tous. Perennials, 20–120 cm, colonial or cespitose; branched rhizomatous, or with branched caudices, becoming ± woody.
Stems

1–3+, ascending to erect (sometimes purple or purplish brown), glabrous or glabrate, sparsely hairy in leaf axils [villous to sericeous].

1–5+, erect (straight to ± flexuous distally, often reddish, sometimes brown), usually glabrous, sometimes ± pilose, particularly distally.

Leaves

thin, apices callous-pointed, faces glabrous or glabrate;

basal usually withering by flowering, long-petiolate, petioles sheathing, sparsely ciliate, blades lanceolate, 40–110 × (3–)5–7 mm, bases attenuate, margins entire to crenulate, rarely with 1–2 teeth, apices obtuse to rounded, callus-pointed;

proximalmost cauline sometimes withering by flowering, winged-petiolate or sessile distally, blades lanceolate to linear-lanceolate, 50–120(–180) × 1–6 mm, bases attenuate, margins entire, sparsely ciliate, apices acute;

distal sessile, blades subulate to lanceolate, 10–50 × 1–2 mm, bases clasping, apices acuminate.

thin, margins serrate (often sharply, teeth acuminate, mucronulate) to serrate-crenate or subentire, strigose, apices acuminate to acute, mucronulate, abaxial faces glabrous or sparsely to densely strigose-pilose, often pilose on midveins, sometimes on other veins also, adaxial glabrous or sparsely to densely strigose, sometimes ± scabrous;

basal withering by flowering, new vernal rosettes often present, long-petiolate (petioles ± narrowly winged, bases dilated, sheathing, ciliate), blades ovate to elliptic or suborbiculate, (10–)35–150 × (10–)25–75 mm, bases usually deeply cordate, sometimes rounded, margins coarsely, often irregularly serrate, apices sometimes obtuse or rounded;

proximal cauline often withering by flowering, winged-petiolate (becoming shorter and more widely winged distally, petiole bases clasping), blades widely to narrowly ovate, 40–100(–140) × 20–40(–70) mm, reduced distally, bases ± deeply cordate to rounded, margins sharply serrate, apices acuminate;

distal usually sessile or subsessile, rarely short-petiolate, blades ovate to lanceolate, 5–105 × 2–45 mm, bases cordate or rounded to attenuate or cuneate, margins serrate or entire (distalmost), apices acuminate.

Peduncles

1–4(–6) cm, ascending, glabrate, bracts 1–4(–5), subulate to lanceolate, foliaceous.

0.3–2 cm, ± pilose, bracts linear-oblanceolate or -lanceolate to linear, foliaceous, distally grading into phyllaries, margins sparsely ciliolate, glabrous.

Involucres

cylindric to hemispheric, (4.5–)5–7.3 mm.

cylindro-campanulate to cylindric, (3–)4.5–5(–6) mm.

Ray florets

14–27;

corollas white, laminae (4.6–)5.6–7(–9.5)–10.3 × 1.3–2 mm.

(8–)10–16(–20);

corollas usually blue to purple, seldom whitish or pink, laminae (5–)6–8(–10) × 1.4–1.8 mm.

Disc florets

18–35(–42);

corollas yellow, 3.5–5 mm, tubes shorter than funnelform throats (hairy), lobes± spreading, triangular, 0.5–0.7 mm.

(8–)10–15(–20);

corollas cream-color or light yellow becoming purple, (3–)4–4.5(–5) mm, tubes slightly shorter than funnelform throats, lobes sometimes ± spreading, narrowly triangular to lanceolate, 0.6–0.9 mm.

Phyllaries

in 2–3(–5) series, subulate to lanceolate, bases indurate, margins hyaline, entire to erose, often ciliate, green zones linear-lanceolate (outer) or rhombic (inner), apices acute, faces glabrous [sparsely villous].

in (3–)4–6 series, linear-lanceolate to oblong-lanceolate, unequal, bases indurate 1/2–3/4, margins scarious, erose, hyaline, sparsely ciliolate, green zones lanceolate to ± diamond-shaped, apical, apices (often red-tipped) acute to obtuse-acuminate or acuminate, mucronulate, faces glabrous or sparsely strigillose.

Heads

(1–)3–20(–40), in paniculiform arrays, branches patent.

[(5–)20–300+] in ± densely paniculiform arrays, branches divaricate to ascending, paniculiform, sometimes ± long-arching, leafy.

Cypselae

light brown to gray, narrowly obovoid to fusiform, sometimes ± compressed, 2–3 mm, 5-nerved, faces glabrate to sparsely strigillose;

pappi white to cream, 3.5–5.5 mm.

dull purple or light brown, obovoid, ± compressed, 2–2.5 mm, 4–5-nerved, faces glabrous;

pappi white or ± rose-tinged, 2.5–4.5 mm.

2n

= 10.

= 16, 32.

Symphyotrichum potosinum

Symphyotrichum cordifolium

Phenology Flowering (Jun–)Jul–Sep. Flowering Aug–Oct.
Habitat Moist, muddy, sandy, or rocky soils along streams Rich, mostly mesic, rocky to loamy soils, open wooded slopes and bluffs, stream banks, moist ledges, swampy woods, border of beech-maple or oak-hickory forets, clearings, thickets, roadsides, along ditches, sometimes weedy in urban areas
Elevation 1500–1900 m (4900–6200 ft) 0–1200 m (0–3900 ft)
Distribution
from FNA
AZ; Mexico
[BONAP county map]
from FNA
AL; AR; CT; DC; GA; IA; IL; IN; KY; MA; MD; ME; MI; MN; MO; NC; NE; NH; NJ; NY; OH; OK; PA; RI; SC; SD; TN; VA; VT; WI; WV; MB; NB; NS; ON; PE; QC
[WildflowerSearch map]
[BONAP county map]
Discussion

Of conservation concern.

Symphyotrichum potosinum is known from Cochise County, Arizona, south into Mexico to northern Oaxaca.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Symphyotrichum cordifolium has been introduced in British Columbia but has not persisted. In a morphometric analysis of infraspecific variation in the northeastern part of the range, A. Legault (1986) showed that the varieties described are not distinct and mostly represent phenotypic variants caused by growing conditions; likewise, diploid and tetraploid races could not be distinguished morphologically.

The type of Aster cordifolius var. laevigatus is conspecific with that of Symphyotrichum cordifolium. This name is the basionym of S. lowrieanum; therefore the latter cannot be considered distinct from S. cordifolium and is not recognized here. Most specimens initially identified as S. lowrieanum in herbaria have been re-determined as S. cordifolium. Some of the specimens, however, appear to correspond to the hybrid between S. cordifolium and S. laeve var. laeve, called S. ×schistosum (Steele) G. L. Nesom (syn. Aster schistosus Steele).

Symphyotrichum ×tardiflorum (Linnaeus) Greuter, Aghababian & Wagenitz [syn. Aster tardiflorus Linnaeus, A. novi-belgii Linnaeus subsp. tardiflorus (Linnaeus) A. G. Jones, A. novi-belgii var. tardiflorus (Linnaeus) A. G. Jones, Symphyotrichum novi-belgii (Linnaeus) G. L. Nesom var. tardiflorum (Linnaeus) G. L. Nesom] is the F1 hybrid between S. cordifolium and S. puniceum (J. Labrecque & Brouillet 1996; G. L. Nesom 1997; W. Greuter et al. 2005). Aster tardiflorus forma vestitus (Fernald) Fernald or var. vestitus Fernald is a hairy variant of the hybrid and is not recognized here; hairiness may be related to whichever species is the maternal parent. A. G. Jones (1989) reported hybrids with S. drummondii and S. urophyllum. She believed that var. moratum may be a hybrid with or an introgressant from S. drummondii, but this is not accepted by all authors. Symphyotrichum ×finkii (Rydberg) G. L. Nesom (syn. Aster finkii Rydberg), may be the hybrid of S. cordifolium and S. shortii. All such putative hybrids need to be confirmed.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 20, p. 480. FNA vol. 20, p. 501.
Parent taxa Asteraceae > tribe Astereae > Symphyotrichum > subg. Astropolium Asteraceae > tribe Astereae > Symphyotrichum > subg. Symphyotrichum > sect. Symphyotrichum
Sibling taxa
S. adnatum, S. anomalum, S. anticostense, S. ascendens, S. boreale, S. bracteolatum, S. campestre, S. chapmanii, S. chilense, S. ciliatum, S. ciliolatum, S. concolor, S. cordifolium, S. cusickii, S. defoliatum, S. depauperatum, S. drummondii, S. dumosum, S. elliottii, S. ericoides, S. eulae, S. falcatum, S. fendleri, S. firmum, S. foliaceum, S. fontinale, S. frondosum, S. georgianum, S. grandiflorum, S. greatae, S. hallii, S. hendersonii, S. jessicae, S. laeve, S. lanceolatum, S. lateriflorum, S. laurentianum, S. lentum, S. molle, S. nahanniense, S. novae-angliae, S. novi-belgii, S. oblongifolium, S. ontarionis, S. oolentangiense, S. parviceps, S. patens, S. phlogifolium, S. pilosum, S. plumosum, S. porteri, S. praealtum, S. pratense, S. prenanthoides, S. priceae, S. puniceum, S. pygmaeum, S. racemosum, S. retroflexum, S. rhiannon, S. robynsianum, S. sericeum, S. shortii, S. simmondsii, S. spathulatum, S. subspicatum, S. subulatum, S. tenuifolium, S. tradescantii, S. turbinellum, S. undulatum, S. urophyllum, S. walteri, S. welshii, S. yukonense, S. ×amethystinum
S. adnatum, S. anomalum, S. anticostense, S. ascendens, S. boreale, S. bracteolatum, S. campestre, S. chapmanii, S. chilense, S. ciliatum, S. ciliolatum, S. concolor, S. cusickii, S. defoliatum, S. depauperatum, S. drummondii, S. dumosum, S. elliottii, S. ericoides, S. eulae, S. falcatum, S. fendleri, S. firmum, S. foliaceum, S. fontinale, S. frondosum, S. georgianum, S. grandiflorum, S. greatae, S. hallii, S. hendersonii, S. jessicae, S. laeve, S. lanceolatum, S. lateriflorum, S. laurentianum, S. lentum, S. molle, S. nahanniense, S. novae-angliae, S. novi-belgii, S. oblongifolium, S. ontarionis, S. oolentangiense, S. parviceps, S. patens, S. phlogifolium, S. pilosum, S. plumosum, S. porteri, S. potosinum, S. praealtum, S. pratense, S. prenanthoides, S. priceae, S. puniceum, S. pygmaeum, S. racemosum, S. retroflexum, S. rhiannon, S. robynsianum, S. sericeum, S. shortii, S. simmondsii, S. spathulatum, S. subspicatum, S. subulatum, S. tenuifolium, S. tradescantii, S. turbinellum, S. undulatum, S. urophyllum, S. walteri, S. welshii, S. yukonense, S. ×amethystinum
Synonyms Aster potosinus, Aster lemmonii Aster cordifolius, Aster cordifolius var. alvearius, Aster cordifolius var. furbishiae, Aster cordifolius var. incisus, Aster cordifolius subsp. laevigatus, Aster cordifolius var. laevigatus, Aster cordifolius var. lanceolatus, Aster cordifolius var. moratus, Aster cordifolius var. polycephalus, Aster cordifolius var. racemiflorus, Aster cordifolius subsp. sagittifolius, Aster cordifolius var. sagittifolius, Aster finkii var. moratus, Aster leiophyllus, Aster lowrieanus, Aster lowrieanus var. incisus, Aster lowrieanus var. lanceolatus, Aster sagittifolius, S. cordifolium var. furbishiae, S. cordifolium var. lanceolatum, S. cordifolium var. moratum, S. cordifolium var. polycephalum, S. cordifolium var. racemiflorum, S. lowrieanum, S. sagittifolium
Name authority (A. Gray) G. L. Nesom: Phytologia 77: 289. (1995) (Linnaeus) G. L. Nesom: Phytologia 77: 278. (1995)
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