Symphyotrichum novae-angliae |
Symphyotrichum anomalum |
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aster de nouvelle-angleterre, New England American-aster, New England aster, New England or michaelmas daisy |
manyray aster |
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Habit | Perennials, 30–120 cm, cespitose; with thick, woody, branched caudices, or short, fleshy rhizomes, sometimes with woody cormoid portions. | Perennials, 20–100(–120) cm, cespitose; usually with short, stout, branched caudices, woody with age, sometimes long-rhizomatous. |
Stems | 1–5+, erect (stout, light to dark brown, sometimes purplish distally), proximally sparsely to moderately hispiduloso-hirsute or pilose, distally moderately to densely so, stipitate-glandular. |
1–5+, erect or ascending (straight, often stout), proximally usually ± densely hirtellous, sometimes glabrescent, distally hirsute. |
Leaves | (light to dark green) thin, often stiff, margins entire or sometimes with shallow teeth, ciliate; basal withered or withering by flowering, sessile, blades (3-nerved) usually spatulate, sometimes oblanceolate, 20–60 × 5–15 mm, bases attenuate, apices acute, faces sparsely hirsute; proximal cauline withering by flowering, sessile, blades oblong or lanceolate, 50–100 × 5–15(–20) mm, bases auriculate-clasping, margins entire, pustulate-scabrous, apices acute, mucronulate, faces stipitate-glandular, abaxial thinly strigose, adaxial hirsute or hispidulous; distal sessile, blades oblanceolate, 30–80 × 6–15 mm, gradually reduced distally, bases auriculate-clasping, apices acute to obtuse, mucronate to minutely white-spinulose, faces moderately to densely short-soft-hairy, sparsely to moderately stipitate-glandular. |
thick, firm, margins (slightly undulate) scabrous, adaxial faces glabrous or strigose, scabrous, adaxial ± sparsely hirsute to scabro-hirtellous, sometimes ± scabrous (cauline sometimes with tufts of smaller leaves in axils); basal withering by flowering, petiolate (petioles sometimes narrowly winged), blades oblong-ovate to lanceolate, 10–40 × 10–20 mm, bases deeply cordate, margins sparsely serrate to serrulate, apices usually acute, sometimes obtuse to rounded; proximal cauline usually withering by flowering, petiolate (petioles slender to ± winged), blades ovate to broadly lanceolate, 40–90 × (15–) 25–50 mm, bases shallowly cordate or rounded to attenuate, rarely truncate, margins subentire or entire (rarely serrulate), apices acute or acuminate, with short callus point; distal subpetiolate or sessile, blades ovate to lanceolate or linear-lanceolate, 10–70 × 1–10 mm, gradually or abruptly reduced distally, bases attenuate to cuneate, margins entire, apices acute to acuminate. |
Peduncles | dilated distally, 0.3–4 cm, densely short-hairy, stipitate-glandular, bracts 1–4, foliaceous, linear to narrowly lanceolate, densely short-hairy, stipitate-glandular, grading into phyllaries. |
densely bracteate, 0.3–5(–10) cm (rarely subsesssile), bracts numerous, linear, 2–6 mm, mucronulate, grading into phyllaries. |
Involucres | campanulate to hemispheric, (5–)7–9(–15) mm. |
campanulate, 5–10 mm. |
Ray florets | (40–)50–75(-100); corollas dark rose to deep purple (pale pink or white), laminae 9–13 × 0.8–1.3 mm. |
18–45; corollas usually bright lavender-blue to purple, seldom white, laminae 9–15(–18) × 1–3.2 mm. |
Disc florets | 50–110; corollas light yellow becoming purple, (4–)4.5–5.5(–7) mm, tubes ± 1/2 narrowly funnelform throats (glabrous or thinly puberulent), lobes triangular, 0.4–0.7 mm. |
33–40+; corollas cream or light yellow turning pinkish purple, (4–)4.5–5.5 mm, tubes shorter than funnelform throats, lobes sometimes ± spreading, triangular to lanceolate, 0.5–0.8 mm. |
Phyllaries | in 3–5(–6) series (dark green to purple-tinged), linear-lanceolate, subequal, outer foliaceous, mid and inner scarious in basal 1/3–1/2, margins stipitate-glandular, apices long-acuminate to acuminate, spreading to reflexed or squarrose, faces glabrous, outer densely stipitate-glandular. |
in 4–6 series, oblong-lanceolate (outer) to oblanceolate-linear (innermost), unequal, bases indurate (appressed), margins ciliate to scabro-ciliate, green zones diamond-shaped to lanceolate, ± foliaceous distally, apices reflexed to squarrose, long-acuminate, apiculate, often purple, faces strigoso-hirtellous. |
Heads | in leafy, often crowded, paniculo-corymbiform arrays. |
in open, diffuse, paniculiform arrays, branches divaricate, stout, long, densely bracteate. |
Cypselae | dull purple or brown, oblong or obconic, not compressed, 1.8–2.5(–3) × 0.6–1 mm, 7–10-nerved, faces densely sericeous, sparsely stipitate-glandular; pappi tawny (barb tips sometimes rose-tinged), 4.5–6 mm. |
deep purple to purplish brown or brown with purple speckles (nerves stramineous), oblong-obovoid, compressed, (1–)2.5–3.5(–3.8) mm, 5–6-nerved, faces glabrous; pappi tawny or rose-tinged, 3.7–5.2 mm. |
2n | = 10. |
= 16. |
Symphyotrichum novae-angliae |
Symphyotrichum anomalum |
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Phenology | Flowering Aug–Oct(–Nov). | Flowering late Aug–Oct. |
Habitat | Open, moist to wet, sandy or loamy, rich soils, fields, prairies, meadows, marshy grounds, shrubby swamps, fens, shores, thickets, moist edges of woods, roadsides, railroad rights-of-way, somewhat weedy | Dry, sandy, loamy, or clayey soils over limestone, or acid soils over chert, sandstone, or granite, rocky, open deciduous woods, thickets, dry ridges, cliffs, bluffs, occasional along streams |
Elevation | 0–1600 m (0–5200 ft) | 50–500 m (200–1600 ft) |
Distribution |
AL; AR; CO; CT; DC; DE; GA; IA; IL; IN; KS; KY; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NY; OH; OK; OR; PA; RI; SC; SD; TN; UT; VA; WA; WI; WV; WY; MB; NB; NS; ON; QC [Introduced in Europe]
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AR; IL; KS; MO; OK
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Discussion | Symphyotrichum novae-angliae is escaped from cultivation and introduced in Montana, Oregon, Utah, Washington, and Wyoming, and has been reported as an ephemeral escape in British Columbia. It possibly escaped from cultivation elsewhere. The Michaelmas daisy is widely sold in the horticultural trade, where cultivars have been developed. Forms have been described that correspond to color genetic variants within natural populations {Aster novae-angliae forma roseus (Desfontaines) Britton; A. novae-angliae forma geneseensis House}; they are not recognized here. Symphyotrichum novae-angliae resembles Canadanthus modestus, but the ranges of the two do not overlap, and the latter has sparsely hairy cypselae with dark ribs. Symphyotrichum novae-angliae hybridizes with S. ericoides, forming the F1 intersectional hybrid S. ×amethystinum. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
A. G. Jones (1989, 1992) reported that Symphyotrichum anomalum may hybridize with S. drummondii, S. oolentangiense, and S. shortii. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 20, p. 487. | FNA vol. 20, p. 504. |
Parent taxa | Asteraceae > tribe Astereae > Symphyotrichum > subg. Virgulus | Asteraceae > tribe Astereae > Symphyotrichum > subg. Symphyotrichum > sect. Symphyotrichum |
Sibling taxa | ||
Synonyms | Aster novae-angliae, Virgulus novae-angliae | Aster anomalus |
Name authority | (Linnaeus) G. L. Nesom: Phytologia 77: 287. (1995) | (Engelmann ex Torrey & A. Gray) G. L. Nesom: Phytologia 77: 275. (1995) |
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