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alpine leafybract aster, Canby's leafybract aster, Cusick's American aster, Cusick's aster, Henderson's aster, Kootenai aster, leafy aster, leafy or leafy-bract or alpine leafybract aster, leafy-bract aster, Parry's aster

aster ciliolé, fringe blue aster, Lindley's American-aster, Lindley's aster, Lindley's or fringe blue aster

Habit Perennials 10–60 cm, colonial; long-rhizomatous. Perennials, (10–)20–120 cm, colonial or cespitose; usually long-rhizomatous, sometimes with branched caudices.
Stems

1–5+, ascending to erect, glabrous or sparsely puberulent.

1–3+, erect (straight to ± flexuous), glabrate to sparsely hirsute or strigillose, especially distally.

Leaves

thin, margins entire or sometimes serrate, apices acute to obtuse, faces usually glabrous, sometimes sparsely hairy;

basal usually persistent, petiolate to subpetiolate, blades broadly elliptic to obovate, 30–200 × 8–25(–30) mm, bases attenuate, margins entire or sometimes serrate, apices acute to obtuse;

proximal cauline sessile or subpetiolate, blades elliptic to obovate, 35–120 × 8–25 mm, bases attenuate or cuneate to rounded, sometimes ± clasping, apices acute;

distal sessile, reduced distally, bases cuneate, apices acute.

thin, margins coarsely, sharply serrate to crenate-serrate or serrulate, ciliate to scabrous, apices acute or acuminate, mucronulate, abaxial faces glabrate to sparsely hirsute, midveins usually densely hirsute, sometimes glabrous, adaxial glabrous or glabrate to scabrellous;

basal usually withering by flowering (sometimes persistent on small plants), long-petiolate (petioles slightly winged, sheathing, ciliate), blades ovate, (24–)40–120(–270) × 15–70 mm, bases usually shallowly cordate, sometimes rounded;

proximal cauline often withering by flowering, winged-petiolate, blades ovate to oblanceolate or lanceolate, (24–)60–150 × (10–)20–60 mm, reduced distally, bases subcordate to cuneate;

distal sessile or sometimes subsessile (petioles widely-winged), blades lance-ovate to linear-lanceolate or linear, 18–75 × (2–)5–25 mm, bases cuneate, sometimes ± clasping, margins serrulate or entire.

Peduncles

sparsely hairy, bracts 0–3, lanceolate.

0.2–1.5+ cm, unequal, reduced distally, ± hirsutulous, bracts 0–4, subulate or linear.

Involucres

campanulate, 6–16(–20) mm.

campanulate, (4–)5–6.5 mm.

Ray florets

15–60;

corollas violet to purple, laminae 8–18(–20) × 1–2 mm.

(12–)14–20;

corollas pale to deep blue or bluish purple, laminae (8.3–)10–15 × 1–2.3 mm.

Disc florets

50–150;

corollas yellow, 4–7 mm, lobes triangular, 0.4–1 mm.

14–25(–29);

corollas yellow becoming reddish purple, 4.3–6.4 mm, tubes shorter than funnelform throats, lobes narrowly triangular, 0.5–1.1 mm.

Phyllaries

in 4–6 series, oblanceolate or oblong (outer) to lanceolate or linear (inner), subequal or unequal (outer exceeding inner), bases outer foliaceous, inner indurate, margins entire, green zones elliptic to lanceolate, apices acute to rounded, faces glabrous or puberulent.

in 4–5 series, narrowly oblong-lanceolate to oblong-oblanceolate or linear-oblanceolate (innermost), ± unequal, bases indurate 1/3–2/3, margins narrowly scarious, erose, hyaline or infrequently purplish, sparsely ciliolate, green zones lanceolate to linear-lanceolate or linear, apices acuminate to long-acuminate, faces glabrous.

Heads

borne singly or in paniculiform arrays, branches ascending, up to 25 cm.

[(6–)13–50(–100+)] in open, paniculiform arrays, branches ascending.

Cypselae

brown, cylindric to obovoid, not compressed, 2.5–4 mm, 3–4-nerved, faces hairy;

pappi white to tawny, 5–8 mm.

yellowish white, obovoid, compressed, 1–2 mm, 5–6-nerved, faces glabrate to sparsely strigillose;

pappi white to pinkish, 3–6 mm.

2n

= 48.

Symphyotrichum foliaceum

Symphyotrichum ciliolatum

Phenology Flowering late Jul–Oct.
Habitat Rich, open, often ± calcareous, boreal deciduous forests (aspen or aspen-birch-fir-spruce), edges of woods, clearings, aspen or bur oak thickets, sometimes open pine forests, streambanks, trails, roadsides
Elevation 0–2000+ m (0–6600+ ft)
Distribution
from FNA
AK; AZ; CA; CO; ID; MT; NM; NV; OR; UT; WA; WY; AB; BC
[WildflowerSearch map]
[BONAP county map]
from FNA
IL; MA; ME; MI; MN; MT; ND; NH; NY; SD; VT; WI; WY; AB; BC; MB; NB; NF; NS; NT; ON; QC; SK; YT
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 4 (4 in the flora).

Symphyotrichum foliaceum is extremely variable and is widespread in western montane coniferous forests and subalpine meadows.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Symphyotrichum ciliolatum sporadically hybridizes with S. laeve var. laeve or var. geyeri in their areas of overlap, notably across the Canadian prairies and in south-central Ontario (Manitoulin Island, Bruce Peninsula). It also hybridizes with S. novi-belgii var. novi-belgii in the Gulf of St. Lawrence area [Symphyotrichum ×subgeminatum (Fernald) G. L. Nesom; syn. Aster subgeminatus (Fernald) B. Boivin]. Aster ciliolatus forma comatus Fernald is densely ciliolate along the midnerves and represents an extreme of the variation found in the species.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Plants 5–30 cm; heads 1–12; phyllaries subequal, narrowly lanceolate
→ 2
1. Plants usually 20–60 cm; heads usually 5–20; outer phyllaries exceeding inner, wider, often foliaceous
→ 3
2. Plants 5–20 cm; heads usually borne singly; phyllaries often purple-tinged; alpine or sub- alpine habitats
var. apricum
2. Plants 10–30 cm; heads usually 1–12; phyllaries green; montane to subalpine habitats.
var. parryi
3. Phyllaries obtuse to rounded; cauline leaves often clasping at base
var. canbyi
3. Phyllaries acute; cauline leaves sometimes rounded at base, not clasping
var. foliaceum
Source FNA vol. 20, p. 532. FNA vol. 20, p. 502.
Parent taxa Asteraceae > tribe Astereae > Symphyotrichum > subg. Symphyotrichum > sect. Occidentales Asteraceae > tribe Astereae > Symphyotrichum > subg. Symphyotrichum > sect. Symphyotrichum
Sibling taxa
S. adnatum, S. anomalum, S. anticostense, S. ascendens, S. boreale, S. bracteolatum, S. campestre, S. chapmanii, S. chilense, S. ciliatum, S. ciliolatum, S. concolor, S. cordifolium, S. cusickii, S. defoliatum, S. depauperatum, S. drummondii, S. dumosum, S. elliottii, S. ericoides, S. eulae, S. falcatum, S. fendleri, S. firmum, S. fontinale, S. frondosum, S. georgianum, S. grandiflorum, S. greatae, S. hallii, S. hendersonii, S. jessicae, S. laeve, S. lanceolatum, S. lateriflorum, S. laurentianum, S. lentum, S. molle, S. nahanniense, S. novae-angliae, S. novi-belgii, S. oblongifolium, S. ontarionis, S. oolentangiense, S. parviceps, S. patens, S. phlogifolium, S. pilosum, S. plumosum, S. porteri, S. potosinum, S. praealtum, S. pratense, S. prenanthoides, S. priceae, S. puniceum, S. pygmaeum, S. racemosum, S. retroflexum, S. rhiannon, S. robynsianum, S. sericeum, S. shortii, S. simmondsii, S. spathulatum, S. subspicatum, S. subulatum, S. tenuifolium, S. tradescantii, S. turbinellum, S. undulatum, S. urophyllum, S. walteri, S. welshii, S. yukonense, S. ×amethystinum
S. adnatum, S. anomalum, S. anticostense, S. ascendens, S. boreale, S. bracteolatum, S. campestre, S. chapmanii, S. chilense, S. ciliatum, S. concolor, S. cordifolium, S. cusickii, S. defoliatum, S. depauperatum, S. drummondii, S. dumosum, S. elliottii, S. ericoides, S. eulae, S. falcatum, S. fendleri, S. firmum, S. foliaceum, S. fontinale, S. frondosum, S. georgianum, S. grandiflorum, S. greatae, S. hallii, S. hendersonii, S. jessicae, S. laeve, S. lanceolatum, S. lateriflorum, S. laurentianum, S. lentum, S. molle, S. nahanniense, S. novae-angliae, S. novi-belgii, S. oblongifolium, S. ontarionis, S. oolentangiense, S. parviceps, S. patens, S. phlogifolium, S. pilosum, S. plumosum, S. porteri, S. potosinum, S. praealtum, S. pratense, S. prenanthoides, S. priceae, S. puniceum, S. pygmaeum, S. racemosum, S. retroflexum, S. rhiannon, S. robynsianum, S. sericeum, S. shortii, S. simmondsii, S. spathulatum, S. subspicatum, S. subulatum, S. tenuifolium, S. tradescantii, S. turbinellum, S. undulatum, S. urophyllum, S. walteri, S. welshii, S. yukonense, S. ×amethystinum
Subordinate taxa
S. foliaceum var. apricum, S. foliaceum var. canbyi, S. foliaceum var. foliaceum, S. foliaceum var. parryi
Synonyms Aster foliaceus Aster ciliolatus, Aster ciliolatus var. comatus, Aster lindleyanus, Aster wilsonii
Name authority (Lindley ex de Candolle) G. L. Nesom: Phytologia 77: 282. (1995) (Lindley) Á. Löve & D. Löve: Taxon 31: 359. (1982)
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