Symphyotrichum ericoides |
Symphyotrichum subg. Virgulus |
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aster éricoïde, heath American-aster, heath-leaf aster, little gray aster, rayless alkali aster, tuft white prairie aster, white heath aster |
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Habit | Perennials, 20–80(–100) cm, colonial or cespitose, eglandular; branched rhizomatous, or with ± cormoid, branched, woody caudices. | Perennials, sometimes stipitate-glandular; rhizomatous or with cormoid caudices. | ||||
Stems | 1–3+, ascending to erect (grayish brown to brown), sparsely to densely hispido-strigose, sometimes glabrescent proximally. |
brittle at maturity, usually ± densely hairy, sometimes glabrous, sometimes stipitate-glandular at least distally. |
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Leaves | usually all except rameal withered by flowering, (light grayish green) firm apices ± white-spine-tipped (often with clusters of smaller leaves in axils); basal sessile, blades (3-nerved) oblanceolate to oblong or spatulate, 10–50 × 10–25 mm, bases attenuate, margins usually entire, rarely remotely serrate, scabrous, apices rounded to obtuse, faces usually sparsely hairy, often glabrous; proximal cauline sessile, blades (1- or 3-nerved) linear to lanceolate or oblong, 10–40(–60) × 1.5–4(–7) mm, reduced distally, bases cuneate, coarsely ciliate, margins entire, coarsely ciliate, apices acute or obtuse, faces moderately to densely strigose or hirsute; distal sessile, blades oblong-ovate, 10–40 × 1.5–3.5 mm, abruptly reduced distally, bases cuneate, margins entire, apices acute, faces moderately to densely strigose. |
basal withering by flowering, usually sessile, sometimes subpetiolate, blades usually 3-nerved, spatulate or obovate to oblanceolate, sometimes linear-oblanceolate, margins entire; cauline sessile, blades ovate or oblong to lanceolate, oblanceolate, or linear, bases rounded or auriculate, clasping or subclasping (leaves sometimes reduced distally). |
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Peduncles | 0.5–1(–2) cm or subsessile, densely hairy, bracts dense, linear to narrowly lanceolate, usually reflexed, sometime appressed to ascending, 1.5–5(–6) mm, densely hairy, grading into phyllaries. |
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Involucres | cylindric to campanulate, 2.5–4.5(–5) mm. |
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Ray florets | (8–)10–18(–20); corollas usually white, rarely pink or bluish, laminae 6–12(–20) × 0.7–1.2 mm. |
7–36(–100) in 1 series; laminae (2–)4.5–24(–25) × 0.5–2 mm. |
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Disc florets/ |
6–12(–20); corollas yellow becoming brown, 2.5–4 mm, throats narrowly funnelform, lobes triangular, 0.5–0.6 mm, glabrous. |
usually weakly ampliate, throats narrowly funnelform, lobes erect. |
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Phyllaries | in 3–4 series, oblanceolate to ± spatulate, unequal, firm, bases (whitish to tan) ± indurate in proximal 1/2–2/3, margins hyaline, scabrous proximally, green zones diamond-shaped, in distal 1/2, apices spine-tipped, (outer) spreading to reflexed or squarrose, faces (outer) sparsely to densely hispid, scabroso-hirsute adaxially, (inner) glabrous. |
unequal or subequal, sometimes ± foliaceous, bases usually indurate, green zones usually diamond-shaped, sometimes lanceolate or apices foliaceous (faces usually hairy, sometimes glabrous, sometimes stipitate-glandular). |
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Heads | (1–200+) in paniculiform arrays, branches fastigiate or arrays often pyramidal, racemiform, secund, crowded. |
radiate. |
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Cypselae | deep purple turning brown, obovoid to oblong-obovoid, ± falcate, not compressed, 1.2–2 × 0.4–0.6 mm, 7–9-nerved (faint), faces sericeous or densely strigillose; pappi whitish, 3–4 mm. |
cylindric to obovoid or oblong, sometimes fusiform, sometimes ± compressed, (5–)6–10-nerved, usually densely strigillose. |
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x | = 5, 4. |
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Symphyotrichum ericoides |
Symphyotrichum subg. Virgulus |
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Distribution |
AR; AZ; CO; CT; DC; DE; IA; ID; IL; IN; KS; MA; MD; ME; MI; MN; MO; MS; MT; ND; NE; NJ; NM; NY; OH; OK; OR; PA; RI; SD; TX; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NT; ON; QC; SK; n Mexico
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North America; Mexico |
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Discussion | Varieties 2 (2 in the flora). Symphyotrichum ericoides resembles S. pilosum var. pilosum, which has larger heads, longer rays, and phyllaries that are not spine-tipped, though the revolute margins can make them appear so. Two subspecies and four weakly separated varieties of S. ericoides were recognized by A. G. Jones (1978). Tetraploids of var. ericoides on the eastern prairies can be difficult to distinguish from S. falcatum. A number of aster cultivars are sold under the name “Aster ericoides.” These are all derived from European garden plants and are either cultivars of S. dumosum, S. lateriflorum, S. pilosum, or S. racemosum, or hybrids involving one of those species and another taxon. The misapplication of the epithet ericoides dates back to the nineteenth century and has persisted in the horticultural literature. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Species 24 (19, including 1 hybrid, in the flora). This group of morphologically and cytologically distinctive asters has been treated at ranks from section to genus. J. C. Semple and L. Brouillet (1980) treated them as the separate genus Lasallea, an illegitimate name replaced by Virgulus (J. L. Reveal and C. S. Keener 1981). A. G. Jones (1980) treated the group as a subgenus within Aster in a broad sense. G. L. Nesom (1994b) proposed subgeneric status for the virguloid asters within Symphyotrichum. On the basis of chloroplast DNA phylogenetic evidence, Xiang C. (1994) showed that the taxon is part of the Symphyotrichum group and recommended treating the virguloid asters as a genus. On the same basis, Xiang and Semple (1996) noted that it could be included within a broadly defined Symphyotrichum or treated separately. Semple et al. (2002), using molecular phylogenetic data, showed the group to belong within Symphyotrichum. Subgenus Virgulus has been divided into four sections. Section Grandiflori (Torrey & A. Gray) G. L. Nesom includes all the stipitate-glandular taxa: S. grandiflorum, S. campestre, S. pygmaeum, S. yukonense, S. fendleri, S. oblongifolium, and S. novae-angliae, sometimes segregated in sect. Polyliguli (Semple & Brouillet) Semple. Section Patentes (Torrey & A. Gray) G. L. Nesom includes S. patens, S. phlogifolium, S. georgianum, S. adnatum, and S. walteri. Section Virgulus (Rafinesque) G. L. Nesom comprises S. concolor, S. plumosum, S. pratense, and S. sericeum. Section Ericoidi (Torrey & A. Gray) G. L. Nesom includes S. ericoides and S. falcatum. Symphyotrichum ×amethystinum is an intersectional hybrid (sects. Grandiflori × Ericoidi) and remains unassigned. Relationships among sections are unresolved. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 20, p. 494. | FNA vol. 20, p. 483. | ||||
Parent taxa | Asteraceae > tribe Astereae > Symphyotrichum > subg. Virgulus | Asteraceae > tribe Astereae > Symphyotrichum | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Aster ericoides, Lasallea ericoides, Virgulus ericoides | subg. Virgulus, Aster subg. Virgulus | ||||
Name authority | (Linnaeus) G. L. Nesom: Phytologia 77: 280. (1995) | (Rafinesque) G. L. Nesom: Phytologia 77: 272. (1995) | ||||
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