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aster éricoïde, heath American-aster, heath-leaf aster, little gray aster, rayless alkali aster, tuft white prairie aster, white heath aster

aster ponceau, purple-stem American-aster, purple-stem aster, purplestem or swamp or red-stem aster, swamp aster

Habit Perennials, 20–80(–100) cm, colonial or cespitose, eglandular; branched rhizomatous, or with ± cormoid, branched, woody caudices. Perennials (7–)100–250(–300) cm, cespitose; thickly short-rhizomatous or with stout, branched caudices, sometimes long-rhizomatous.
Stems

1–3+, ascending to erect (grayish brown to brown), sparsely to densely hispido-strigose, sometimes glabrescent proximally.

1–5+, erect (straight to distally flexuous, stout 5–11 mm diam., usually reddish purple), usually densely to sparsely hirsute or hispid, glabrescent, often proximally glabrate, sometimes glabrate.

Leaves

usually all except rameal withered by flowering, (light grayish green) firm apices ± white-spine-tipped (often with clusters of smaller leaves in axils);

basal sessile, blades (3-nerved) oblanceolate to oblong or spatulate, 10–50 × 10–25 mm, bases attenuate, margins usually entire, rarely remotely serrate, scabrous, apices rounded to obtuse, faces usually sparsely hairy, often glabrous;

proximal cauline sessile, blades (1- or 3-nerved) linear to lanceolate or oblong, 10–40(–60) × 1.5–4(–7) mm, reduced distally, bases cuneate, coarsely ciliate, margins entire, coarsely ciliate, apices acute or obtuse, faces moderately to densely strigose or hirsute;

distal sessile, blades oblong-ovate, 10–40 × 1.5–3.5 mm, abruptly reduced distally, bases cuneate, margins entire, apices acute, faces moderately to densely strigose.

(dark green, not crowded) thin, margins remotely serrate to sometimes entire, scabrous, apices acute or acuminate to subcaudate, mucronate, abaxial faces glabrate, ± densely pilose along midveins, adaxial glabrate to scabrous;

basal withering by flowering, subpetiolate to petiolate (petioles ± winged, bases dilated, sheathing), blades spatulate to oblanceolate or lanceolate, 30–10+ × 3–20+ mm, bases attenuate to cuneate, margins remotely crenate-serrate to denticulate or entire, apices acute or acuminate to rounded;

proximal cauline withering by flowering, petiolate to subpetiolate (petioles widely winged, bases auriculate, clasping), blades lanceolate or oblanceolate to oblong, elliptic or elliptic-oblong, (55–)70–200(–220) × 10–40 mm, usually progressively lengthened to mid, bases ± attenuate to cuneate or auriculate and clasping;

distal sessile, blades oblanceolate or lanceolate to oblong or elliptic-oblong (sometimes nearly panduriform), 8–160 × 1–40 mm, progressively reduced distally, strongly so and linear in arrays, bases ± strongly auriculate, clasping, margins subentire or entire.

Peduncles

0.5–1(–2) cm or subsessile, densely hairy, bracts dense, linear to narrowly lanceolate, usually reflexed, sometime appressed to ascending, 1.5–5(–6) mm, densely hairy, grading into phyllaries.

0.2–3+ cm, ± hispid or hirsute to villous, bracts 0–3, lanceolate or linear-lanceolate to linear, distal often subtending heads.

Involucres

cylindric to campanulate, 2.5–4.5(–5) mm.

campanulate, (6–)8–12(–15) mm.

Ray florets

(8–)10–18(–20);

corollas usually white, rarely pink or bluish, laminae 6–12(–20) × 0.7–1.2 mm.

20–50(–60);

corollas usually pale to dark blue, lavender or purple, seldom white or pink, laminae (7–)12–18(–21) × (0.9–)1.4–1.8 mm.

Disc florets

6–12(–20);

corollas yellow becoming brown, 2.5–4 mm, throats narrowly funnelform, lobes triangular, 0.5–0.6 mm, glabrous.

30–50 (–90);

corollas yellow or cream becoming pink or purple, (4.1–)5–6.5 mm, tubes shorter than cylindro-funnelform limbs, lobes triangular, 0.4–1.1 mm.

Phyllaries

in 3–4 series, oblanceolate to ± spatulate, unequal, firm, bases (whitish to tan) ± indurate in proximal 1/2–2/3, margins hyaline, scabrous proximally, green zones diamond-shaped, in distal 1/2, apices spine-tipped, (outer) spreading to reflexed or squarrose, faces (outer) sparsely to densely hispid, scabroso-hirsute adaxially, (inner) glabrous.

in 4–6 series, oblanceolate-spatulate to linear, subequal, bases indurate 1/5–1/2, margins not scarious (outer) to narrowly scarious, erose, hyaline, sparsely ciliolate, green zones linear-lanceolate, outer sometimes ± foliaceous, to linear (inner), apices ± spreading at tip when foliaceous, long-acuminate to caudate, mucronate to apiculate, sometimes purplish tinged, faces glabrous or ± hispid.

Heads

(1–200+) in paniculiform arrays, branches fastigiate or arrays often pyramidal, racemiform, secund, crowded.

in open, paniculiform (often corymbiform-looking) arrays, branches widely spreading to ascending, sparsely to moderately leafy with ± large leaves.

Cypselae

deep purple turning brown, obovoid to oblong-obovoid, ± falcate, not compressed, 1.2–2 × 0.4–0.6 mm, 7–9-nerved (faint), faces sericeous or densely strigillose;

pappi whitish, 3–4 mm.

purple or brown (nerves stramineous), oblanceoloid, ± falcate, compressed, (2–)2.5–3.5(–4) mm, 3–4-nerved (nerves prominent), faces glabrous or sparsely strigillose;

pappi white, 3.8–6 mm.

Symphyotrichum ericoides

Symphyotrichum puniceum

Distribution
from FNA
AR; AZ; CO; CT; DC; DE; IA; ID; IL; IN; KS; MA; MD; ME; MI; MN; MO; MS; MT; ND; NE; NJ; NM; NY; OH; OK; OR; PA; RI; SD; TX; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NT; ON; QC; SK; n Mexico
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; CT; DE; GA; IA; IL; IN; LA; MD; ME; MI; MN; MO; MS; NC; ND; NE; NH; NJ; NY; OH; PA; RI; SC; SD; TN; TX; VA; VT; WI; WV; AB; BC; MB; NB; NS; ON; PE; QC; SK
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

Symphyotrichum ericoides resembles S. pilosum var. pilosum, which has larger heads, longer rays, and phyllaries that are not spine-tipped, though the revolute margins can make them appear so. Two subspecies and four weakly separated varieties of S. ericoides were recognized by A. G. Jones (1978). Tetraploids of var. ericoides on the eastern prairies can be difficult to distinguish from S. falcatum. A number of aster cultivars are sold under the name “Aster ericoides.” These are all derived from European garden plants and are either cultivars of S. dumosum, S. lateriflorum, S. pilosum, or S. racemosum, or hybrids involving one of those species and another taxon. The misapplication of the epithet ericoides dates back to the nineteenth century and has persisted in the horticultural literature.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

Symphyotrichum puniceum is one of the widest ranging species in the genus, growing from the edges of the prairies to the Atlantic coast, and from the Gulf coast of Texas to southern Ungava Bay (northern Quebec).

Several taxa have been recognized within this variable species. Here, we segregate Symphyotrichum firmum as a distinct entity. The tall, pink-rayed S. elliottii of the outer coastal plains is also distinctive, despite reports of hybrids with S. puniceum var. puniceum where their ranges overlap (see G. L. Nesom 1997b). A recently described member of the complex, S. rhiannon, is an ultramafic endemic from North Carolina. Here, within S. puniceum, we recognize two varieties, the variable and widespread var. puniceum, and var. scabricaule from the inner Gulf coastal plains, following Nesom, who also offered a useful summary of the infraspecific taxonomy of the species.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Plants colonial, rhizomatous, not cormoid; involucres cylindro-campanulate when fresh
var. ericoides
1. Plants cespitose, with cormoid caudices, not strongly rhizomatous; involucres broadly campanulate when fresh
var. pansum
1. Stems sparsely to ± densely hirsute, glabrescent, sometimes glabrate proximally; abaxial leaf faces paler than adaxial, with a dark, distinct reticulum, adaxial without impressed main veins; array leaves usually equal to mid cauline
var. puniceum
1. Stems densely and uniformly hirsute; abaxial and adaxial leaf faces ±concolorous, without dark, distinct reticulum, adaxial with distinctly impressed main veins; array leaves reduced in size relative to mid cauline
var. scabricaule
Source FNA vol. 20, p. 494. FNA vol. 20, p. 524.
Parent taxa Asteraceae > tribe Astereae > Symphyotrichum > subg. Virgulus Asteraceae > tribe Astereae > Symphyotrichum > subg. Symphyotrichum > sect. Symphyotrichum
Sibling taxa
S. adnatum, S. anomalum, S. anticostense, S. ascendens, S. boreale, S. bracteolatum, S. campestre, S. chapmanii, S. chilense, S. ciliatum, S. ciliolatum, S. concolor, S. cordifolium, S. cusickii, S. defoliatum, S. depauperatum, S. drummondii, S. dumosum, S. elliottii, S. eulae, S. falcatum, S. fendleri, S. firmum, S. foliaceum, S. fontinale, S. frondosum, S. georgianum, S. grandiflorum, S. greatae, S. hallii, S. hendersonii, S. jessicae, S. laeve, S. lanceolatum, S. lateriflorum, S. laurentianum, S. lentum, S. molle, S. nahanniense, S. novae-angliae, S. novi-belgii, S. oblongifolium, S. ontarionis, S. oolentangiense, S. parviceps, S. patens, S. phlogifolium, S. pilosum, S. plumosum, S. porteri, S. potosinum, S. praealtum, S. pratense, S. prenanthoides, S. priceae, S. puniceum, S. pygmaeum, S. racemosum, S. retroflexum, S. rhiannon, S. robynsianum, S. sericeum, S. shortii, S. simmondsii, S. spathulatum, S. subspicatum, S. subulatum, S. tenuifolium, S. tradescantii, S. turbinellum, S. undulatum, S. urophyllum, S. walteri, S. welshii, S. yukonense, S. ×amethystinum
S. adnatum, S. anomalum, S. anticostense, S. ascendens, S. boreale, S. bracteolatum, S. campestre, S. chapmanii, S. chilense, S. ciliatum, S. ciliolatum, S. concolor, S. cordifolium, S. cusickii, S. defoliatum, S. depauperatum, S. drummondii, S. dumosum, S. elliottii, S. ericoides, S. eulae, S. falcatum, S. fendleri, S. firmum, S. foliaceum, S. fontinale, S. frondosum, S. georgianum, S. grandiflorum, S. greatae, S. hallii, S. hendersonii, S. jessicae, S. laeve, S. lanceolatum, S. lateriflorum, S. laurentianum, S. lentum, S. molle, S. nahanniense, S. novae-angliae, S. novi-belgii, S. oblongifolium, S. ontarionis, S. oolentangiense, S. parviceps, S. patens, S. phlogifolium, S. pilosum, S. plumosum, S. porteri, S. potosinum, S. praealtum, S. pratense, S. prenanthoides, S. priceae, S. pygmaeum, S. racemosum, S. retroflexum, S. rhiannon, S. robynsianum, S. sericeum, S. shortii, S. simmondsii, S. spathulatum, S. subspicatum, S. subulatum, S. tenuifolium, S. tradescantii, S. turbinellum, S. undulatum, S. urophyllum, S. walteri, S. welshii, S. yukonense, S. ×amethystinum
Subordinate taxa
S. ericoides var. ericoides, S. ericoides var. pansum
S. puniceum var. puniceum, S. puniceum var. scabricaule
Synonyms Aster ericoides, Lasallea ericoides, Virgulus ericoides Aster puniceus
Name authority (Linnaeus) G. L. Nesom: Phytologia 77: 280. (1995) (Linnaeus) Á. Löve & D. Löve: Taxon 31: 359. (1982)
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