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aster éricoïde, heath American-aster, heath-leaf aster, little gray aster, rayless alkali aster, tuft white prairie aster, white heath aster

aster cordifolié, common blue wood aster, heart-leaf American-aster, heart-leaf aster, heartleaf or common blue wood aster

Habit Perennials, 20–80(–100) cm, colonial or cespitose, eglandular; branched rhizomatous, or with ± cormoid, branched, woody caudices. Perennials, 20–120 cm, colonial or cespitose; branched rhizomatous, or with branched caudices, becoming ± woody.
Stems

1–3+, ascending to erect (grayish brown to brown), sparsely to densely hispido-strigose, sometimes glabrescent proximally.

1–5+, erect (straight to ± flexuous distally, often reddish, sometimes brown), usually glabrous, sometimes ± pilose, particularly distally.

Leaves

usually all except rameal withered by flowering, (light grayish green) firm apices ± white-spine-tipped (often with clusters of smaller leaves in axils);

basal sessile, blades (3-nerved) oblanceolate to oblong or spatulate, 10–50 × 10–25 mm, bases attenuate, margins usually entire, rarely remotely serrate, scabrous, apices rounded to obtuse, faces usually sparsely hairy, often glabrous;

proximal cauline sessile, blades (1- or 3-nerved) linear to lanceolate or oblong, 10–40(–60) × 1.5–4(–7) mm, reduced distally, bases cuneate, coarsely ciliate, margins entire, coarsely ciliate, apices acute or obtuse, faces moderately to densely strigose or hirsute;

distal sessile, blades oblong-ovate, 10–40 × 1.5–3.5 mm, abruptly reduced distally, bases cuneate, margins entire, apices acute, faces moderately to densely strigose.

thin, margins serrate (often sharply, teeth acuminate, mucronulate) to serrate-crenate or subentire, strigose, apices acuminate to acute, mucronulate, abaxial faces glabrous or sparsely to densely strigose-pilose, often pilose on midveins, sometimes on other veins also, adaxial glabrous or sparsely to densely strigose, sometimes ± scabrous;

basal withering by flowering, new vernal rosettes often present, long-petiolate (petioles ± narrowly winged, bases dilated, sheathing, ciliate), blades ovate to elliptic or suborbiculate, (10–)35–150 × (10–)25–75 mm, bases usually deeply cordate, sometimes rounded, margins coarsely, often irregularly serrate, apices sometimes obtuse or rounded;

proximal cauline often withering by flowering, winged-petiolate (becoming shorter and more widely winged distally, petiole bases clasping), blades widely to narrowly ovate, 40–100(–140) × 20–40(–70) mm, reduced distally, bases ± deeply cordate to rounded, margins sharply serrate, apices acuminate;

distal usually sessile or subsessile, rarely short-petiolate, blades ovate to lanceolate, 5–105 × 2–45 mm, bases cordate or rounded to attenuate or cuneate, margins serrate or entire (distalmost), apices acuminate.

Peduncles

0.5–1(–2) cm or subsessile, densely hairy, bracts dense, linear to narrowly lanceolate, usually reflexed, sometime appressed to ascending, 1.5–5(–6) mm, densely hairy, grading into phyllaries.

0.3–2 cm, ± pilose, bracts linear-oblanceolate or -lanceolate to linear, foliaceous, distally grading into phyllaries, margins sparsely ciliolate, glabrous.

Involucres

cylindric to campanulate, 2.5–4.5(–5) mm.

cylindro-campanulate to cylindric, (3–)4.5–5(–6) mm.

Ray florets

(8–)10–18(–20);

corollas usually white, rarely pink or bluish, laminae 6–12(–20) × 0.7–1.2 mm.

(8–)10–16(–20);

corollas usually blue to purple, seldom whitish or pink, laminae (5–)6–8(–10) × 1.4–1.8 mm.

Disc florets

6–12(–20);

corollas yellow becoming brown, 2.5–4 mm, throats narrowly funnelform, lobes triangular, 0.5–0.6 mm, glabrous.

(8–)10–15(–20);

corollas cream-color or light yellow becoming purple, (3–)4–4.5(–5) mm, tubes slightly shorter than funnelform throats, lobes sometimes ± spreading, narrowly triangular to lanceolate, 0.6–0.9 mm.

Phyllaries

in 3–4 series, oblanceolate to ± spatulate, unequal, firm, bases (whitish to tan) ± indurate in proximal 1/2–2/3, margins hyaline, scabrous proximally, green zones diamond-shaped, in distal 1/2, apices spine-tipped, (outer) spreading to reflexed or squarrose, faces (outer) sparsely to densely hispid, scabroso-hirsute adaxially, (inner) glabrous.

in (3–)4–6 series, linear-lanceolate to oblong-lanceolate, unequal, bases indurate 1/2–3/4, margins scarious, erose, hyaline, sparsely ciliolate, green zones lanceolate to ± diamond-shaped, apical, apices (often red-tipped) acute to obtuse-acuminate or acuminate, mucronulate, faces glabrous or sparsely strigillose.

Heads

(1–200+) in paniculiform arrays, branches fastigiate or arrays often pyramidal, racemiform, secund, crowded.

[(5–)20–300+] in ± densely paniculiform arrays, branches divaricate to ascending, paniculiform, sometimes ± long-arching, leafy.

Cypselae

deep purple turning brown, obovoid to oblong-obovoid, ± falcate, not compressed, 1.2–2 × 0.4–0.6 mm, 7–9-nerved (faint), faces sericeous or densely strigillose;

pappi whitish, 3–4 mm.

dull purple or light brown, obovoid, ± compressed, 2–2.5 mm, 4–5-nerved, faces glabrous;

pappi white or ± rose-tinged, 2.5–4.5 mm.

2n

= 16, 32.

Symphyotrichum ericoides

Symphyotrichum cordifolium

Phenology Flowering Aug–Oct.
Habitat Rich, mostly mesic, rocky to loamy soils, open wooded slopes and bluffs, stream banks, moist ledges, swampy woods, border of beech-maple or oak-hickory forets, clearings, thickets, roadsides, along ditches, sometimes weedy in urban areas
Elevation 0–1200 m (0–3900 ft)
Distribution
from FNA
AR; AZ; CO; CT; DC; DE; IA; ID; IL; IN; KS; MA; MD; ME; MI; MN; MO; MS; MT; ND; NE; NJ; NM; NY; OH; OK; OR; PA; RI; SD; TX; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NT; ON; QC; SK; n Mexico
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; CT; DC; GA; IA; IL; IN; KY; MA; MD; ME; MI; MN; MO; NC; NE; NH; NJ; NY; OH; OK; PA; RI; SC; SD; TN; VA; VT; WI; WV; MB; NB; NS; ON; PE; QC
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

Symphyotrichum ericoides resembles S. pilosum var. pilosum, which has larger heads, longer rays, and phyllaries that are not spine-tipped, though the revolute margins can make them appear so. Two subspecies and four weakly separated varieties of S. ericoides were recognized by A. G. Jones (1978). Tetraploids of var. ericoides on the eastern prairies can be difficult to distinguish from S. falcatum. A number of aster cultivars are sold under the name “Aster ericoides.” These are all derived from European garden plants and are either cultivars of S. dumosum, S. lateriflorum, S. pilosum, or S. racemosum, or hybrids involving one of those species and another taxon. The misapplication of the epithet ericoides dates back to the nineteenth century and has persisted in the horticultural literature.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Symphyotrichum cordifolium has been introduced in British Columbia but has not persisted. In a morphometric analysis of infraspecific variation in the northeastern part of the range, A. Legault (1986) showed that the varieties described are not distinct and mostly represent phenotypic variants caused by growing conditions; likewise, diploid and tetraploid races could not be distinguished morphologically.

The type of Aster cordifolius var. laevigatus is conspecific with that of Symphyotrichum cordifolium. This name is the basionym of S. lowrieanum; therefore the latter cannot be considered distinct from S. cordifolium and is not recognized here. Most specimens initially identified as S. lowrieanum in herbaria have been re-determined as S. cordifolium. Some of the specimens, however, appear to correspond to the hybrid between S. cordifolium and S. laeve var. laeve, called S. ×schistosum (Steele) G. L. Nesom (syn. Aster schistosus Steele).

Symphyotrichum ×tardiflorum (Linnaeus) Greuter, Aghababian & Wagenitz [syn. Aster tardiflorus Linnaeus, A. novi-belgii Linnaeus subsp. tardiflorus (Linnaeus) A. G. Jones, A. novi-belgii var. tardiflorus (Linnaeus) A. G. Jones, Symphyotrichum novi-belgii (Linnaeus) G. L. Nesom var. tardiflorum (Linnaeus) G. L. Nesom] is the F1 hybrid between S. cordifolium and S. puniceum (J. Labrecque & Brouillet 1996; G. L. Nesom 1997; W. Greuter et al. 2005). Aster tardiflorus forma vestitus (Fernald) Fernald or var. vestitus Fernald is a hairy variant of the hybrid and is not recognized here; hairiness may be related to whichever species is the maternal parent. A. G. Jones (1989) reported hybrids with S. drummondii and S. urophyllum. She believed that var. moratum may be a hybrid with or an introgressant from S. drummondii, but this is not accepted by all authors. Symphyotrichum ×finkii (Rydberg) G. L. Nesom (syn. Aster finkii Rydberg), may be the hybrid of S. cordifolium and S. shortii. All such putative hybrids need to be confirmed.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Plants colonial, rhizomatous, not cormoid; involucres cylindro-campanulate when fresh
var. ericoides
1. Plants cespitose, with cormoid caudices, not strongly rhizomatous; involucres broadly campanulate when fresh
var. pansum
Source FNA vol. 20, p. 494. FNA vol. 20, p. 501.
Parent taxa Asteraceae > tribe Astereae > Symphyotrichum > subg. Virgulus Asteraceae > tribe Astereae > Symphyotrichum > subg. Symphyotrichum > sect. Symphyotrichum
Sibling taxa
S. adnatum, S. anomalum, S. anticostense, S. ascendens, S. boreale, S. bracteolatum, S. campestre, S. chapmanii, S. chilense, S. ciliatum, S. ciliolatum, S. concolor, S. cordifolium, S. cusickii, S. defoliatum, S. depauperatum, S. drummondii, S. dumosum, S. elliottii, S. eulae, S. falcatum, S. fendleri, S. firmum, S. foliaceum, S. fontinale, S. frondosum, S. georgianum, S. grandiflorum, S. greatae, S. hallii, S. hendersonii, S. jessicae, S. laeve, S. lanceolatum, S. lateriflorum, S. laurentianum, S. lentum, S. molle, S. nahanniense, S. novae-angliae, S. novi-belgii, S. oblongifolium, S. ontarionis, S. oolentangiense, S. parviceps, S. patens, S. phlogifolium, S. pilosum, S. plumosum, S. porteri, S. potosinum, S. praealtum, S. pratense, S. prenanthoides, S. priceae, S. puniceum, S. pygmaeum, S. racemosum, S. retroflexum, S. rhiannon, S. robynsianum, S. sericeum, S. shortii, S. simmondsii, S. spathulatum, S. subspicatum, S. subulatum, S. tenuifolium, S. tradescantii, S. turbinellum, S. undulatum, S. urophyllum, S. walteri, S. welshii, S. yukonense, S. ×amethystinum
S. adnatum, S. anomalum, S. anticostense, S. ascendens, S. boreale, S. bracteolatum, S. campestre, S. chapmanii, S. chilense, S. ciliatum, S. ciliolatum, S. concolor, S. cusickii, S. defoliatum, S. depauperatum, S. drummondii, S. dumosum, S. elliottii, S. ericoides, S. eulae, S. falcatum, S. fendleri, S. firmum, S. foliaceum, S. fontinale, S. frondosum, S. georgianum, S. grandiflorum, S. greatae, S. hallii, S. hendersonii, S. jessicae, S. laeve, S. lanceolatum, S. lateriflorum, S. laurentianum, S. lentum, S. molle, S. nahanniense, S. novae-angliae, S. novi-belgii, S. oblongifolium, S. ontarionis, S. oolentangiense, S. parviceps, S. patens, S. phlogifolium, S. pilosum, S. plumosum, S. porteri, S. potosinum, S. praealtum, S. pratense, S. prenanthoides, S. priceae, S. puniceum, S. pygmaeum, S. racemosum, S. retroflexum, S. rhiannon, S. robynsianum, S. sericeum, S. shortii, S. simmondsii, S. spathulatum, S. subspicatum, S. subulatum, S. tenuifolium, S. tradescantii, S. turbinellum, S. undulatum, S. urophyllum, S. walteri, S. welshii, S. yukonense, S. ×amethystinum
Subordinate taxa
S. ericoides var. ericoides, S. ericoides var. pansum
Synonyms Aster ericoides, Lasallea ericoides, Virgulus ericoides Aster cordifolius, Aster cordifolius var. alvearius, Aster cordifolius var. furbishiae, Aster cordifolius var. incisus, Aster cordifolius subsp. laevigatus, Aster cordifolius var. laevigatus, Aster cordifolius var. lanceolatus, Aster cordifolius var. moratus, Aster cordifolius var. polycephalus, Aster cordifolius var. racemiflorus, Aster cordifolius subsp. sagittifolius, Aster cordifolius var. sagittifolius, Aster finkii var. moratus, Aster leiophyllus, Aster lowrieanus, Aster lowrieanus var. incisus, Aster lowrieanus var. lanceolatus, Aster sagittifolius, S. cordifolium var. furbishiae, S. cordifolium var. lanceolatum, S. cordifolium var. moratum, S. cordifolium var. polycephalum, S. cordifolium var. racemiflorum, S. lowrieanum, S. sagittifolium
Name authority (Linnaeus) G. L. Nesom: Phytologia 77: 280. (1995) (Linnaeus) G. L. Nesom: Phytologia 77: 278. (1995)
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