The green links below add additional plants to the comparison table. Blue links lead to other Web sites.
enable glossary links

bushy American-aster, bushy aster, rice button aster

annual saltmarsh American-aster, annual saltmarsh aster, annual saltmarsh or eastern annual saltmarsh aster, aster subulé, eastern annual saltmarsh aster, Southwestern annual saltmarsh aster

Habit Perennials, (20–)30–100 cm, colonial or cespitose; usually long-rhizomatous, often thick, woody, sometimes short, or with short, stout caudices. Annuals, (10–)30–150 cm; tap-rooted.
Stems

1–5+, erect (straight, often slender, brittle), densely or sparsely strigose to glabrescent or glabrate.

1, erect (often with purple or purplish brown areas), glabrous or glabrate, sometimes strigillose in leaf axils.

Leaves

firm, margins entire to crenulate-serrate, recurved, scabrous, apices sharply white-mucronate, abaxial faces glabrous or glabrate to sparsely strigillose, adaxial glabrous or glabrate to submarginally scabrous (short-strigose), cauline withering by flowering (except on array branches), often with axillary leaf clusters;

basal withering by flowering, petiolate, petioles winged, sheathing, strigose-ciliate, blades oblanceolate to spatulate (often declined), 8–50 × 3–15 mm, bases attenuate to cuneate, margins crenate-serrate, apices obtuse or rounded to ± acute;

proximal cauline (mostly declined) subpetiolate (petioles widely winged, clasping) or sessile (then not or barely clasping), blades linear-oblanceolate, 25–120 × 1.5–9 mm, bases cuneate to slightly attenuate;

distal (ascending or spreading) sessile, blades oblong or linear-lanceolate to linear, 2–45 × 0.5–4 mm, reduced distally (abruptly on branches), bases cuneate to rounded, not clasping, margins entire or serrate.

thin (green to dark green), margins often strigilloso-ciliolate, faces glabrous;

basal withering by flowering, long-petiolate (petiole bases sheathing), sparsely ciliate, blades ovate to oblanceolate, 10–90 × 6–14 mm, bases attenuate to cuneate, rounded, margins entire or serrulate or crenulate, apices rounded, obtuse, or acute;

proximal cauline withering by flowering, petiolate, subpetiolate, or sessile, blades narrowly lanceolate or subulate, 20–100(–200) × 1.5–10(–20) mm, bases attenuate, margins subentire, entire, or serrulate, apices acute to acuminate;

distal sessile, blades narrowly lanceolate to subulate, 5–113 × 0.5–5.5 mm, apices acuminate.

Peduncles

slender, usually stiff (sometimes lax), (0.5–)1–5 cm (rarely subsessile), progressively reduced distally, usually not secund, sparsely strigillose or glabrous, bracts 5–16+, spreading to ascending, linear-oblong or -elliptic to subulate or linear, progressively reduced distally, grading into phyllaries.

(0.2–)0.5–4 cm, bracts 4–8(–17).

Involucres

cylindro-campanulate, (3–)4.5–6.3 mm.

cylindric to turbinate, 5–7(–8.2) mm.

Ray florets

15–33;

corollas pale blue, pink, or lavender to white, laminae (4–)5–7(–8) × 1–1.7 mm.

16–30(–54) in 1–3 series;

corollas white, pink, or lavender, laminae 1.3–7 × 0.2–1.3 mm.

Disc florets

15–30;

corollas cream to pale yellow turning pink, 3.5–4.5 mm, tubes shorter than to equal to narrowly funnelform throats, lobes lanceolate, (0.4–)0.6–1.1 mm.

4–10(–13);

corollas yello, sometimes tinged with purple, 3.4–5.2 mm, throats narrowly funnelform, lobes ± spreading to erect, narrowly triangular, 0.3–0.7 mm, glabrous.

Phyllaries

in 4–6 series, appressed or slightly spreading, oblong-oblanceolate (outer) to linear-oblanceolate or linear (innermost), strongly unequal, bases indurate (1/2–)2/3–4/5, margins hyaline, scarious, erose, distally ciliolate, green zones oblanceolate to elliptic (subapical), apices acute to obtuse (scarious), sometimes faintly reddish, mucronulate, faces usually glabrous, sometimes glabrate.

in 3–5 series, broadly or narrowly lanceolate to subulate, unequal, bases indurate, margins hyaline, often purple-tinged, entire, green zones lanceolate (usually narrow, sometimes broad and covering most of distal portion), apices acute, faces glabrous.

Heads

in remote, diffuse, open, paniculiform arrays, branches numerous, ascending to stiffly ascending, secondary ones stiff, racemiform (usually not secund).

(10–) 30–100(–150), in open, diffuse, paniculiform arrays.

Cypselae

pink or stramineous with pink streaks, or gray (± dark, nerves stramineous), oblong-obovoid, sometimes ± compressed, 1.5–2.5 mm, 3–4-nerved (nerves prominent), faces strigillose;

pappi white to sordid, 4 mm.

light brown to purple, narrowly obovoid to fusiform, sometimes ± compressed, (1.2–)1.5–2.7(–3) mm, 5-nerved, faces sparsely strigillose;

pappi white, (3–)3.5–5.5 mm.

2n

= 16, 32.

Symphyotrichum dumosum

Symphyotrichum subulatum

Phenology Flowering Aug–Oct.
Habitat Open or wooded, moist or wet soils, bogs, fens, sedge meadows, marshes, swamps, flood plains, sandy or calcareous flats, loamy prairies, old fields, sandhills, flatwoods, hammocks, pine-hickory woods, oak or pine thickets, secondary woods, sandy to mucky or marly shores of lakes and ponds, interdunal hollows
Elevation 0–700 m (0–2300 ft)
Distribution
from FNA
AL; AR; CT; DC; DE; FL; GA; IA; IL; IN; KY; LA; MA; MD; ME; MI; MO; MS; NC; NH; NJ; NY; OH; OK; PA; RI; SC; TN; TX; VA; WI; WV; NB; ON
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; AZ; CA; CT; DE; FL; GA; IL; IN; KS; LA; MA; MD; ME; MI; MS; NC; NE; NH; NJ; NM; NV; NY; OH; OK; PA; RI; SC; TN; TX; UT; VA; NB; ON; Mexico; Central America; South America; West Indies; Bermuda [Widely introduced worldwide]
[WildflowerSearch map]
[BONAP county map]
Discussion

Symphyotrichum dumosum is introduced in New Brunswick. It is widely cultivated. It can hybridize with S. racemosum and S. lanceolatum var. interior (A. G. Jones 1989). G. L. Nesom (1994b) and J. C. Semple et al. (2002) recognized several varieties within the complex: var. dumosum [syn. Aster coridifolius Michaux, A. dumosus Linnaeus var. coridifolius (Michaux) Torrey & A. Gray]; var. gracilipes (Wiegand) G. L. Nesom (syn. A. dumosus var. gracilipes Wiegand); var. pergracile (Wiegand) G. L. Nesom (syn. A. dumosus var. pergracile Wiegand); var. strictior (Torrey & A. Gray) G. L. Nesom [syn. A. dumosus var. strictior Torrey & A. Gray, A. dumosus var. dodgei Fernald, S. dumosum var. dodgei (Fernald) G. L. Nesom, the latter possibly the hybrid S. dumosum × S. boreale fide G. L. Nesom 1997]; and var. subulifolium (Torrey & A. Gray) G. L. Nesom (syn. A. dumosus var. subulifolium Torrey & A. Gray). Some of these taxa may deserve higher rank. More work is needed in the complex, however, before a coherent taxonomy can be achieved.

G. L. Nesom (1997) recognized also a southern pine flatwoods entity called Symphyotrichum kralii G. L. Nesom, based on the illegitimately named Aster pinifolius Alexander (not Nees, nor F. Mueller). The type of A. pinifolius, however, appears to be conspecific with that of S. simmondsii. At present, it seems best to wait for full investigation before formally recognizing such taxa.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 5 (5 in the flora).

Five varieties of Symphyotrichum subulatum are recognized for North America based on differences in chromosome number, ray lamina color and size, array shapes, number of series of ray florets, number of disc and ray florets, and other, more cryptic characters (S. D. Sundberg 2004). These varieties were treated as species by G. L. Nesom (1994b, 2005d). Variety ligulatum is apparently an obligate outcrosser and is the least variable variety (Sundberg). Other varieties are self-compatible, which could facilitate the fixation of mutations in populations.

The five varieties are nearly entirely allopatric, and intermediates between pairs of varieties are not uncommon where they approach one another. Populations that are intermediate in ray lamina size between vars. ligulatum and parviflorum are widespread in southern Texas, New Mexico, Arizona, and northern Mexico. Intermediates between vars. elongatum and parviflorum and between vars. elongatum and subulatum occur in Florida. Despite these observations, hybridization experiments and chromosome number differences suggest that the varieties are mostly reproductively isolated (S. D. Sundberg 1986, 2004).

In older floras the name Aster exilis Elliott has been applied to Symphyotrichum subulatum vars. ligulatum and parviflorum. The status of this name is uncertain; the type specimen has been lost and the description of the plant is inadequate for determining the taxon to which the name should be applied (G. L. Nesom 1994b; S. D. Sundberg 2004).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Ray laminae lavender to blue, (3.5–)4.5–7 × 0.9–1.3 mm, drying in 3–5 coils; ray florets in 1 series; disc florets (20–)33–50 (sc United States)
var. ligulatum
1. Ray laminae white to pink or lavender, 1.3–3.5(–4.2) × 0.2–0.6 mm, drying in (0–)1–3(–4) coils; ray florets in 1–3 series; disc florets 3–23
→ 2
2. Ray laminae pink to lavender, (2–)2.5–3.5(–4.2) × 0.3–0.6 mm, drying in 2–3(–4) coils; disc florets 11–23 (Florida, West Indies)
var. elongatum
2. Ray laminae usually white, rarely pink or lavender, 1.3–3 × 0.2–0.5 mm, drying in 1 coil or curling; disc florets (3–)4–15
→ 3
3. Ray laminae longer than pappi; phyllaries 30–42; pappi 3.5–3.8(–4.2) mm; w United States
var. parviflorum
3. Ray laminae shorter to slightly longer than pappi; phyllaries 18–30; pappi 3.5–5.5 mm; e and se United States
→ 4
4. Green zones of phyllaries narrowly to broadly lanceolate, extending phyllary length; ray laminae 0.2–0.5 mm wide; disc florets 3.8–4.6(–4.9) mm; salt marshes, e North America and Gulf Coast
var. subulatum
4. Green zones of phyllaries broadly lanceolate, nearly absent from proximal portion; ray laminae 0.2–0.3 mm wide; disc florets 3.2–4.1 mm; adventive, s United States
var. squamatum
Source FNA vol. 20, p. 514. FNA vol. 20, p. 480.
Parent taxa Asteraceae > tribe Astereae > Symphyotrichum > subg. Symphyotrichum > sect. Symphyotrichum Asteraceae > tribe Astereae > Symphyotrichum > subg. Astropolium
Sibling taxa
S. adnatum, S. anomalum, S. anticostense, S. ascendens, S. boreale, S. bracteolatum, S. campestre, S. chapmanii, S. chilense, S. ciliatum, S. ciliolatum, S. concolor, S. cordifolium, S. cusickii, S. defoliatum, S. depauperatum, S. drummondii, S. elliottii, S. ericoides, S. eulae, S. falcatum, S. fendleri, S. firmum, S. foliaceum, S. fontinale, S. frondosum, S. georgianum, S. grandiflorum, S. greatae, S. hallii, S. hendersonii, S. jessicae, S. laeve, S. lanceolatum, S. lateriflorum, S. laurentianum, S. lentum, S. molle, S. nahanniense, S. novae-angliae, S. novi-belgii, S. oblongifolium, S. ontarionis, S. oolentangiense, S. parviceps, S. patens, S. phlogifolium, S. pilosum, S. plumosum, S. porteri, S. potosinum, S. praealtum, S. pratense, S. prenanthoides, S. priceae, S. puniceum, S. pygmaeum, S. racemosum, S. retroflexum, S. rhiannon, S. robynsianum, S. sericeum, S. shortii, S. simmondsii, S. spathulatum, S. subspicatum, S. subulatum, S. tenuifolium, S. tradescantii, S. turbinellum, S. undulatum, S. urophyllum, S. walteri, S. welshii, S. yukonense, S. ×amethystinum
S. adnatum, S. anomalum, S. anticostense, S. ascendens, S. boreale, S. bracteolatum, S. campestre, S. chapmanii, S. chilense, S. ciliatum, S. ciliolatum, S. concolor, S. cordifolium, S. cusickii, S. defoliatum, S. depauperatum, S. drummondii, S. dumosum, S. elliottii, S. ericoides, S. eulae, S. falcatum, S. fendleri, S. firmum, S. foliaceum, S. fontinale, S. frondosum, S. georgianum, S. grandiflorum, S. greatae, S. hallii, S. hendersonii, S. jessicae, S. laeve, S. lanceolatum, S. lateriflorum, S. laurentianum, S. lentum, S. molle, S. nahanniense, S. novae-angliae, S. novi-belgii, S. oblongifolium, S. ontarionis, S. oolentangiense, S. parviceps, S. patens, S. phlogifolium, S. pilosum, S. plumosum, S. porteri, S. potosinum, S. praealtum, S. pratense, S. prenanthoides, S. priceae, S. puniceum, S. pygmaeum, S. racemosum, S. retroflexum, S. rhiannon, S. robynsianum, S. sericeum, S. shortii, S. simmondsii, S. spathulatum, S. subspicatum, S. tenuifolium, S. tradescantii, S. turbinellum, S. undulatum, S. urophyllum, S. walteri, S. welshii, S. yukonense, S. ×amethystinum
Subordinate taxa
S. subulatum var. elongatum, S. subulatum var. ligulatum, S. subulatum var. parviflorum, S. subulatum var. squamatum, S. subulatum var. subulatum
Synonyms Aster dumosus, Aster coridifolius Aster subulatus
Name authority (Linnaeus) G. L. Nesom: Phytologia 77: 280. (1995) (Michaux) G. L. Nesom: Phytologia 77: 293. (1995)
Web links