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Drummond's aster

Anticosti American-aster, Anticosti aster, Anticosti Island aster, aster d'Anticosti

Habit Perennials, 30–120 cm, cespitose; short-rhizomatous or with stout, often woody caudices. Perennials, 10–90 cm, colonial; long-rhizomatous.
Stems

1–5+, erect (straight, stout), glabrate or glabrescent, sparsely pilose, or hirsuto-pilose proximally, ± densely so distally.

1, erect (straight, often reddish), glabrous.

Leaves

(abaxially pale or grayish green, adaxially dark green) initially ± firm or membranous, becoming thicker, margins crenate-serrate to serrate, strigoso-ciliate to scabrous, apices acuminate to acute, mucronate, abaxial faces scabrous to densely piloso-strigose, more so on veins, adaxial scabrous to glabrate;

basal withering by flowering, long-petiolate (petioles narrowly winged, sheathing, sometimes ciliate), blades ovate to ovate-oblong or -lanceolate to elliptic or suborbiculate, 10–85(–100) × 8–60 mm, bases deeply cordate to rounded or abruptly attenuate, margins crenate-serrate, apices usually rounded to obtuse, rarely acute;

proximal cauline often withering by flowering, long-petiolate (petioles narrowly winged, more broadly so distally, ciliate), blades ovate or ovate-lanceolate, (25–)50–120(–150) × (15–)20–50(–65) mm, progressively to slightly reduced distally, bases deeply to shallowly cordate or rounded, margins crenate-serrate to sharply serrate;

distal petiolate or subpetiolate or sessile, petioles broadly winged, blades ovate to ovate- or oblong-lanceolate, to oblong-elliptic or linear-lanceolate (4–)5–110 × 1–50 mm, reduced distally (sharply so on array branches in var. texanum), bases shallowly cordate or rounded to attenuate or cuneate (distally), margins crenate-serrate or -serrulate or entire.

coriaceous, margins remotely serrulate or entire, slightly revolute, scabrous, apices mucronulate, faces glabrous;

basal withering by flowering, long-petiolate (petioles narrowly winged, bases sheathing), blades lanceolate to oblanceolate, ca. 55+ × 3+ mm, bases slightly attenuate, apices acute;

proximal cauline mostly withering by flowering, petiolate (petioles narrowly winged, sheathing), blades lanceolate or oblanceolate to linear-lanceolate or -oblanceolate, arcuate, 90–160 × 5–18 mm, bases cuneate to slightly attenuate, margins entire or serrulate, apices acute;

distal sessile, blades linear to linear-lanceolate, 8–50 × 1.5–5 mm, progressively reduced distally, more strongly so in arrays, bases cuneate to rounded, sometimes slightly clasping, apices acute to acuminate.

Peduncles

0, or 0.2–1.5(–4) cm, densely ascending- or appressed-bracteate, pilosulous, bracts 4–17+, lanceolate or subulate to linear-oblanceolate, distally grading into phyllaries, pilosulous or strigoso-ciliate.

0.8–5.7 cm, slender, sparsely ot moderately pilosulous, bracts 3–5, linear-lanceolate, often crowded proximal to and surpassing heads.

Involucres

campanulate, (3.5–)4.5–6(–6.5) mm.

campanulate, 6–10 mm.

Ray florets

(8–)10–15(—20);

corollas usually bright blue to purple or lavender, sometimes white, laminae (5–)7–10(–12) × 1–1.8 mm.

25–44;

corollas usually pale purple or lilac, sometimes white, laminae 9.5–20 × 0.7–1.4 mm.

Disc florets

(10–)13–15(–22);

corollas cream to yellow, becoming deep purple or reddish purple, 3.5–4.5(–5) mm, tubes shorter than funnelform to cylindric throats, lobes triangular to lanceolate, 0.4–0.7 mm.

29–52;

corollas yellow becoming reddish purple, 5–6 mm, tubes slightly shorter than funnelform throats, lobes lanceolate, 0.6–0.7 mm.

Phyllaries

in 4–5 series, subulate or lanceolate (outermost) to narrowly oblong-lanceolate, unequal, bases indurate 1/4–3/4, margins narrowly scarious proximally, hyaline, ciliate, bodies sometimes green to margins distally, green zones lanceolate to linear-lanceolate, apices acute to acuminate or long-acuminate, tips often purple, sometimes involute, mucronulate, faces glabrous.

in 2–3 series, oblong-oblanceolate or -spatulate (outer) or oblong-lanceolate to linear-lanceolate or linear (inner), ± unequal, bases indurate 1/3–3/4, margins hyaline, scarious, erose distally, sparsely ciliolate distally, green zones lanceolate, outer often foliaceous distally, apices acute to long-acuminate, ± spreading, faces glabrous.

Heads

in open, paniculiform arrays, branches ascending to divaricate, often densely racemiform or ± paniculiform (lateral ones open, racemiform) to subsecund, ± stiff to lax, long to relatively short, ± closely to ± remotely leafy, branch leaves progressively or ± abruptly reduced distally.

in elongate, loosely racemiform arrays, branches ascending, heads single at ends of primary branches or long pedicels.

Cypselae

dull purple or brown to tan (ribs tan to stramineous), obovoid, ± compressed, (1.5–)2–3 mm, 5–6-nerved, faces glabrate to sparsely strigillose distally;

pappi cream to rose-tinged, 3–4 mm.

reddish tan, obovoid, compressed, 1.5–2.8 mm, 4–5-nerved, faces strigillose;

pappi yellowish, 7–8 mm.

2n

= 80.

Symphyotrichum drummondii

Symphyotrichum anticostense

Phenology Flowering Aug–Sep.
Habitat Calcareous, gravelly shores of rivers, lakeshore limestone pavements
Elevation 0–100 m (0–300 ft)
Distribution
from FNA
AL; AR; GA; IA; IL; IN; KS; KY; LA; MD; MI; MN; MO; MS; NE; OH; OK; PA; TN; TX; WI; WV; ne Mexico
[WildflowerSearch map]
[BONAP county map]
from FNA
ME; NB; QC
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

Variety texanum is tetraploid; var. drummondii is both diploid and tetraploid. The tetraploid race of var. drummondii is most likely autopolyploid; origin of var. texanum has not been determined. The two taxa intergrade and do not appear to warrant species status.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

Symphyotrichum anticostense is known from disjunct areas: western Lake St. John, southern Anticosti Island, southern streams of the Gaspé Peninsula (Quebec), Restigouche River (Quebec–New Brunswick), and St. John (New Brunswick) and Aroostook (Maine; A. Haines 2000) rivers. It is an allopolyploid derivative of the cross between the calcareous fen species S. boreale and the widespread shore species S. novi-belgii (J. Labrecque and L. Brouillet 1990). It hybridizes with S. novi-belgii.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Plants 40–120 cm; stems proximally sparsely hirsute, distally densely so; leaves firm, basal and proximal long-petiolate, petioles ± winged, proximal bases ± cordate or truncate, sometimes obliquely; heads in ample, paniculiform arrays with divaricate or ascending, bracteate, often racemiform branches; peduncles ± secund, 0–4 cm, bracteate; involucres campanulate, (3.5–)4.5–7 mm; ray corollas usually bright blue, light purple, bluish violet, or lavender, sometimes white; cypselae glabrous
var. drummondii
1. Plants 30–80 cm; stems ± densely hirsute, particularly distally; leaves membranous, becoming thickish, brittle, basal and proximal winged-petiolate, proximal bases cordate, becoming rounded-truncate distally; heads in open, paniculiform arrays with very long, widely spreading branches; peduncles 1–4 cm, densely, minutely bracteate; involucres turbinate to hemispheric, 3.8–5.2 mm; ray corollas bluish white; cypselae strigillose
var. texanum
Source FNA vol. 20, p. 503. FNA vol. 20, p. 529.
Parent taxa Asteraceae > tribe Astereae > Symphyotrichum > subg. Symphyotrichum > sect. Symphyotrichum Asteraceae > tribe Astereae > Symphyotrichum > subg. Symphyotrichum > sect. Symphyotrichum
Sibling taxa
S. adnatum, S. anomalum, S. anticostense, S. ascendens, S. boreale, S. bracteolatum, S. campestre, S. chapmanii, S. chilense, S. ciliatum, S. ciliolatum, S. concolor, S. cordifolium, S. cusickii, S. defoliatum, S. depauperatum, S. dumosum, S. elliottii, S. ericoides, S. eulae, S. falcatum, S. fendleri, S. firmum, S. foliaceum, S. fontinale, S. frondosum, S. georgianum, S. grandiflorum, S. greatae, S. hallii, S. hendersonii, S. jessicae, S. laeve, S. lanceolatum, S. lateriflorum, S. laurentianum, S. lentum, S. molle, S. nahanniense, S. novae-angliae, S. novi-belgii, S. oblongifolium, S. ontarionis, S. oolentangiense, S. parviceps, S. patens, S. phlogifolium, S. pilosum, S. plumosum, S. porteri, S. potosinum, S. praealtum, S. pratense, S. prenanthoides, S. priceae, S. puniceum, S. pygmaeum, S. racemosum, S. retroflexum, S. rhiannon, S. robynsianum, S. sericeum, S. shortii, S. simmondsii, S. spathulatum, S. subspicatum, S. subulatum, S. tenuifolium, S. tradescantii, S. turbinellum, S. undulatum, S. urophyllum, S. walteri, S. welshii, S. yukonense, S. ×amethystinum
S. adnatum, S. anomalum, S. ascendens, S. boreale, S. bracteolatum, S. campestre, S. chapmanii, S. chilense, S. ciliatum, S. ciliolatum, S. concolor, S. cordifolium, S. cusickii, S. defoliatum, S. depauperatum, S. drummondii, S. dumosum, S. elliottii, S. ericoides, S. eulae, S. falcatum, S. fendleri, S. firmum, S. foliaceum, S. fontinale, S. frondosum, S. georgianum, S. grandiflorum, S. greatae, S. hallii, S. hendersonii, S. jessicae, S. laeve, S. lanceolatum, S. lateriflorum, S. laurentianum, S. lentum, S. molle, S. nahanniense, S. novae-angliae, S. novi-belgii, S. oblongifolium, S. ontarionis, S. oolentangiense, S. parviceps, S. patens, S. phlogifolium, S. pilosum, S. plumosum, S. porteri, S. potosinum, S. praealtum, S. pratense, S. prenanthoides, S. priceae, S. puniceum, S. pygmaeum, S. racemosum, S. retroflexum, S. rhiannon, S. robynsianum, S. sericeum, S. shortii, S. simmondsii, S. spathulatum, S. subspicatum, S. subulatum, S. tenuifolium, S. tradescantii, S. turbinellum, S. undulatum, S. urophyllum, S. walteri, S. welshii, S. yukonense, S. ×amethystinum
Subordinate taxa
S. drummondii var. drummondii, S. drummondii var. texanum
Synonyms Aster drummondii, Aster sagittifolius var. drummondii Aster anticostensis, Aster gaspensis, Aster hesperius var. gaspensis
Name authority (Lindley) G. L. Nesom: Phytologia 77: 279. (1995) (Fernald) G. L. Nesom: Phytologia 77: 275. (1995)
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