Symphyotrichum cordifolium |
Symphyotrichum ontarionis |
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aster cordifolié, common blue wood aster, heart-leaf American-aster, heart-leaf aster, heartleaf or common blue wood aster |
aster du lac Ontario, bottomland aster, Ontario American-aster, Ontario aster |
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Habit | Perennials, 20–120 cm, colonial or cespitose; branched rhizomatous, or with branched caudices, becoming ± woody. | Perennials, 20–120 cm, colonial; long-rhizomatous. | ||||
Stems | 1–5+, erect (straight to ± flexuous distally, often reddish, sometimes brown), usually glabrous, sometimes ± pilose, particularly distally. |
1–(2–3), ascending to erect (straight), proximally glabrate, distally uniformly villous or hirsute, or glabrous (var. glabratum). |
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Leaves | thin, margins serrate (often sharply, teeth acuminate, mucronulate) to serrate-crenate or subentire, strigose, apices acuminate to acute, mucronulate, abaxial faces glabrous or sparsely to densely strigose-pilose, often pilose on midveins, sometimes on other veins also, adaxial glabrous or sparsely to densely strigose, sometimes ± scabrous; basal withering by flowering, new vernal rosettes often present, long-petiolate (petioles ± narrowly winged, bases dilated, sheathing, ciliate), blades ovate to elliptic or suborbiculate, (10–)35–150 × (10–)25–75 mm, bases usually deeply cordate, sometimes rounded, margins coarsely, often irregularly serrate, apices sometimes obtuse or rounded; proximal cauline often withering by flowering, winged-petiolate (becoming shorter and more widely winged distally, petiole bases clasping), blades widely to narrowly ovate, 40–100(–140) × 20–40(–70) mm, reduced distally, bases ± deeply cordate to rounded, margins sharply serrate, apices acuminate; distal usually sessile or subsessile, rarely short-petiolate, blades ovate to lanceolate, 5–105 × 2–45 mm, bases cordate or rounded to attenuate or cuneate, margins serrate or entire (distalmost), apices acuminate. |
thin, margins scabrous, apices mucronate, abaxial faces usually sparsely to densely strigose or strigillose, sometimes glabrous (var. glabrum), adaxial usually strigose or scabrous, sometimes glabrate or glabrous (var. glabrum); basal withering by flowering, petiolate to subpetiolate (petioles narrowly winged, ciliate, bases sheathing), blades spatulate to oblanceolate-obovate, 10–40 × 5–10 mm, bases attenuate, margins crenate-serrate, apices acute to rounded; proximalmost cauline usually withering by flowering, petiolate or sessile (petioles narrowly winged, ± clasping), blades ovate or lance-ovate to elliptic-lanceolate or oblanceolate, 20–80(–12) × 5–35 mm, progressively reduced distally, bases attenuate to cuneate, margins serrate (sometimes coarsely) to crenate-serrate, apices acute to acuminate or short-caudate; distal sessile, blades elliptic-lanceolate to oblanceolate or lanceolate, (6–)10–80+ × 2–25 mm, progressively reduced distally, bases cuneate, margins entire to serrulate, apices acute to acuminate. |
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Peduncles | 0.3–2 cm, ± pilose, bracts linear-oblanceolate or -lanceolate to linear, foliaceous, distally grading into phyllaries, margins sparsely ciliolate, glabrous. |
subsessile or 2–10(–20) mm, ± pilose, bracts 1–5, linear-lanceolate, pilose, grading into phyllaries. |
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Involucres | cylindro-campanulate to cylindric, (3–)4.5–5(–6) mm. |
campanulate, 3–5.5 mm. |
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Ray florets | (8–)10–16(–20); corollas usually blue to purple, seldom whitish or pink, laminae (5–)6–8(–10) × 1.4–1.8 mm. |
(10–)15–26; corollas usually white, rarely pinkish or light purple to blue, laminae 3.5–5.5(–8) × 0.5–1.5 mm. |
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Disc florets | (8–)10–15(–20); corollas cream-color or light yellow becoming purple, (3–)4–4.5(–5) mm, tubes slightly shorter than funnelform throats, lobes sometimes ± spreading, narrowly triangular to lanceolate, 0.6–0.9 mm. |
12–25; corollas cream or light yellow turning magenta or purple (ampliate), 2.5–4(–4.5) mm, tubes shorter than funnelform throats, lobes slightly spreading to reflexed, lanceolate, 0.7–1.1 mm. |
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Phyllaries | in (3–)4–6 series, linear-lanceolate to oblong-lanceolate, unequal, bases indurate 1/2–3/4, margins scarious, erose, hyaline, sparsely ciliolate, green zones lanceolate to ± diamond-shaped, apical, apices (often red-tipped) acute to obtuse-acuminate or acuminate, mucronulate, faces glabrous or sparsely strigillose. |
in (3–)4–6 series, unequal, appressed or ± spreading, linear-obovate (outer) to oblong-lanceolate to linear (inner), bases indurate 1/3–3/4, margins narrowly scarious to apices, erose, hyaline, ciliate, green zones lanceolate, apices acute to acuminate, mucronulate, faces (outer) sparsely pilose or glabrous (var. glabratum), (inner) glabrous. |
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Heads | [(5–)20–300+] in ± densely paniculiform arrays, branches divaricate to ascending, paniculiform, sometimes ± long-arching, leafy. |
in ample, open, paniculiform arrays, branches ± ascending or divaricate to long-arching, ± secund. |
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Cypselae | dull purple or light brown, obovoid, ± compressed, 2–2.5 mm, 4–5-nerved, faces glabrous; pappi white or ± rose-tinged, 2.5–4.5 mm. |
gray or tan, oblong-obovoid, sometimes ± compressed, 1.2–1.8(–2) mm, 3–5-nerved, faces strigillose; pappi whitish to white, 3–3.5 mm. |
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2n | = 16, 32. |
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Symphyotrichum cordifolium |
Symphyotrichum ontarionis |
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Phenology | Flowering Aug–Oct. | |||||
Habitat | Rich, mostly mesic, rocky to loamy soils, open wooded slopes and bluffs, stream banks, moist ledges, swampy woods, border of beech-maple or oak-hickory forets, clearings, thickets, roadsides, along ditches, sometimes weedy in urban areas | |||||
Elevation | 0–1200 m (0–3900 ft) | |||||
Distribution |
AL; AR; CT; DC; GA; IA; IL; IN; KY; MA; MD; ME; MI; MN; MO; NC; NE; NH; NJ; NY; OH; OK; PA; RI; SC; SD; TN; VA; VT; WI; WV; MB; NB; NS; ON; PE; QC
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AL; IA; IL; IN; KS; KY; LA; MI; MN; MO; MS; NC; NE; NY; OK; PA; SD; TN; TX; VT; WI; ON; QC
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Discussion | Symphyotrichum cordifolium has been introduced in British Columbia but has not persisted. In a morphometric analysis of infraspecific variation in the northeastern part of the range, A. Legault (1986) showed that the varieties described are not distinct and mostly represent phenotypic variants caused by growing conditions; likewise, diploid and tetraploid races could not be distinguished morphologically. The type of Aster cordifolius var. laevigatus is conspecific with that of Symphyotrichum cordifolium. This name is the basionym of S. lowrieanum; therefore the latter cannot be considered distinct from S. cordifolium and is not recognized here. Most specimens initially identified as S. lowrieanum in herbaria have been re-determined as S. cordifolium. Some of the specimens, however, appear to correspond to the hybrid between S. cordifolium and S. laeve var. laeve, called S. ×schistosum (Steele) G. L. Nesom (syn. Aster schistosus Steele). Symphyotrichum ×tardiflorum (Linnaeus) Greuter, Aghababian & Wagenitz [syn. Aster tardiflorus Linnaeus, A. novi-belgii Linnaeus subsp. tardiflorus (Linnaeus) A. G. Jones, A. novi-belgii var. tardiflorus (Linnaeus) A. G. Jones, Symphyotrichum novi-belgii (Linnaeus) G. L. Nesom var. tardiflorum (Linnaeus) G. L. Nesom] is the F1 hybrid between S. cordifolium and S. puniceum (J. Labrecque & Brouillet 1996; G. L. Nesom 1997; W. Greuter et al. 2005). Aster tardiflorus forma vestitus (Fernald) Fernald or var. vestitus Fernald is a hairy variant of the hybrid and is not recognized here; hairiness may be related to whichever species is the maternal parent. A. G. Jones (1989) reported hybrids with S. drummondii and S. urophyllum. She believed that var. moratum may be a hybrid with or an introgressant from S. drummondii, but this is not accepted by all authors. Symphyotrichum ×finkii (Rydberg) G. L. Nesom (syn. Aster finkii Rydberg), may be the hybrid of S. cordifolium and S. shortii. All such putative hybrids need to be confirmed. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 2 (2 in the flora). Symphyotrichum ontarionis is often confused with S. lateriflorum, from which it can be distinguished by its shorter disc corolla lobes and abaxial leaf faces either moderately to densely hairy or glabrous to glabrate (var. glabratum), and without hairs along midveins (as is typical of S. lateriflorum). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 20, p. 501. | FNA vol. 20, p. 518. | ||||
Parent taxa | ||||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Aster cordifolius, Aster cordifolius var. alvearius, Aster cordifolius var. furbishiae, Aster cordifolius var. incisus, Aster cordifolius subsp. laevigatus, Aster cordifolius var. laevigatus, Aster cordifolius var. lanceolatus, Aster cordifolius var. moratus, Aster cordifolius var. polycephalus, Aster cordifolius var. racemiflorus, Aster cordifolius subsp. sagittifolius, Aster cordifolius var. sagittifolius, Aster finkii var. moratus, Aster leiophyllus, Aster lowrieanus, Aster lowrieanus var. incisus, Aster lowrieanus var. lanceolatus, Aster sagittifolius, S. cordifolium var. furbishiae, S. cordifolium var. lanceolatum, S. cordifolium var. moratum, S. cordifolium var. polycephalum, S. cordifolium var. racemiflorum, S. lowrieanum, S. sagittifolium | Aster ontarionis, Aster missouriensis | ||||
Name authority | (Linnaeus) G. L. Nesom: Phytologia 77: 278. (1995) | (Wiegand) G. L. Nesom: Phytologia 77: 287. (1995) | ||||
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