Symphyotrichum cordifolium |
Symphyotrichum novae-angliae |
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aster cordifolié, common blue wood aster, heart-leaf American-aster, heart-leaf aster, heartleaf or common blue wood aster |
aster de nouvelle-angleterre, New England American-aster, New England aster, New England or michaelmas daisy |
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Habit | Perennials, 20–120 cm, colonial or cespitose; branched rhizomatous, or with branched caudices, becoming ± woody. | Perennials, 30–120 cm, cespitose; with thick, woody, branched caudices, or short, fleshy rhizomes, sometimes with woody cormoid portions. |
Stems | 1–5+, erect (straight to ± flexuous distally, often reddish, sometimes brown), usually glabrous, sometimes ± pilose, particularly distally. |
1–5+, erect (stout, light to dark brown, sometimes purplish distally), proximally sparsely to moderately hispiduloso-hirsute or pilose, distally moderately to densely so, stipitate-glandular. |
Leaves | thin, margins serrate (often sharply, teeth acuminate, mucronulate) to serrate-crenate or subentire, strigose, apices acuminate to acute, mucronulate, abaxial faces glabrous or sparsely to densely strigose-pilose, often pilose on midveins, sometimes on other veins also, adaxial glabrous or sparsely to densely strigose, sometimes ± scabrous; basal withering by flowering, new vernal rosettes often present, long-petiolate (petioles ± narrowly winged, bases dilated, sheathing, ciliate), blades ovate to elliptic or suborbiculate, (10–)35–150 × (10–)25–75 mm, bases usually deeply cordate, sometimes rounded, margins coarsely, often irregularly serrate, apices sometimes obtuse or rounded; proximal cauline often withering by flowering, winged-petiolate (becoming shorter and more widely winged distally, petiole bases clasping), blades widely to narrowly ovate, 40–100(–140) × 20–40(–70) mm, reduced distally, bases ± deeply cordate to rounded, margins sharply serrate, apices acuminate; distal usually sessile or subsessile, rarely short-petiolate, blades ovate to lanceolate, 5–105 × 2–45 mm, bases cordate or rounded to attenuate or cuneate, margins serrate or entire (distalmost), apices acuminate. |
(light to dark green) thin, often stiff, margins entire or sometimes with shallow teeth, ciliate; basal withered or withering by flowering, sessile, blades (3-nerved) usually spatulate, sometimes oblanceolate, 20–60 × 5–15 mm, bases attenuate, apices acute, faces sparsely hirsute; proximal cauline withering by flowering, sessile, blades oblong or lanceolate, 50–100 × 5–15(–20) mm, bases auriculate-clasping, margins entire, pustulate-scabrous, apices acute, mucronulate, faces stipitate-glandular, abaxial thinly strigose, adaxial hirsute or hispidulous; distal sessile, blades oblanceolate, 30–80 × 6–15 mm, gradually reduced distally, bases auriculate-clasping, apices acute to obtuse, mucronate to minutely white-spinulose, faces moderately to densely short-soft-hairy, sparsely to moderately stipitate-glandular. |
Peduncles | 0.3–2 cm, ± pilose, bracts linear-oblanceolate or -lanceolate to linear, foliaceous, distally grading into phyllaries, margins sparsely ciliolate, glabrous. |
dilated distally, 0.3–4 cm, densely short-hairy, stipitate-glandular, bracts 1–4, foliaceous, linear to narrowly lanceolate, densely short-hairy, stipitate-glandular, grading into phyllaries. |
Involucres | cylindro-campanulate to cylindric, (3–)4.5–5(–6) mm. |
campanulate to hemispheric, (5–)7–9(–15) mm. |
Ray florets | (8–)10–16(–20); corollas usually blue to purple, seldom whitish or pink, laminae (5–)6–8(–10) × 1.4–1.8 mm. |
(40–)50–75(-100); corollas dark rose to deep purple (pale pink or white), laminae 9–13 × 0.8–1.3 mm. |
Disc florets | (8–)10–15(–20); corollas cream-color or light yellow becoming purple, (3–)4–4.5(–5) mm, tubes slightly shorter than funnelform throats, lobes sometimes ± spreading, narrowly triangular to lanceolate, 0.6–0.9 mm. |
50–110; corollas light yellow becoming purple, (4–)4.5–5.5(–7) mm, tubes ± 1/2 narrowly funnelform throats (glabrous or thinly puberulent), lobes triangular, 0.4–0.7 mm. |
Phyllaries | in (3–)4–6 series, linear-lanceolate to oblong-lanceolate, unequal, bases indurate 1/2–3/4, margins scarious, erose, hyaline, sparsely ciliolate, green zones lanceolate to ± diamond-shaped, apical, apices (often red-tipped) acute to obtuse-acuminate or acuminate, mucronulate, faces glabrous or sparsely strigillose. |
in 3–5(–6) series (dark green to purple-tinged), linear-lanceolate, subequal, outer foliaceous, mid and inner scarious in basal 1/3–1/2, margins stipitate-glandular, apices long-acuminate to acuminate, spreading to reflexed or squarrose, faces glabrous, outer densely stipitate-glandular. |
Heads | [(5–)20–300+] in ± densely paniculiform arrays, branches divaricate to ascending, paniculiform, sometimes ± long-arching, leafy. |
in leafy, often crowded, paniculo-corymbiform arrays. |
Cypselae | dull purple or light brown, obovoid, ± compressed, 2–2.5 mm, 4–5-nerved, faces glabrous; pappi white or ± rose-tinged, 2.5–4.5 mm. |
dull purple or brown, oblong or obconic, not compressed, 1.8–2.5(–3) × 0.6–1 mm, 7–10-nerved, faces densely sericeous, sparsely stipitate-glandular; pappi tawny (barb tips sometimes rose-tinged), 4.5–6 mm. |
2n | = 16, 32. |
= 10. |
Symphyotrichum cordifolium |
Symphyotrichum novae-angliae |
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Phenology | Flowering Aug–Oct. | Flowering Aug–Oct(–Nov). |
Habitat | Rich, mostly mesic, rocky to loamy soils, open wooded slopes and bluffs, stream banks, moist ledges, swampy woods, border of beech-maple or oak-hickory forets, clearings, thickets, roadsides, along ditches, sometimes weedy in urban areas | Open, moist to wet, sandy or loamy, rich soils, fields, prairies, meadows, marshy grounds, shrubby swamps, fens, shores, thickets, moist edges of woods, roadsides, railroad rights-of-way, somewhat weedy |
Elevation | 0–1200 m (0–3900 ft) | 0–1600 m (0–5200 ft) |
Distribution |
AL; AR; CT; DC; GA; IA; IL; IN; KY; MA; MD; ME; MI; MN; MO; NC; NE; NH; NJ; NY; OH; OK; PA; RI; SC; SD; TN; VA; VT; WI; WV; MB; NB; NS; ON; PE; QC
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AL; AR; CO; CT; DC; DE; GA; IA; IL; IN; KS; KY; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NY; OH; OK; OR; PA; RI; SC; SD; TN; UT; VA; WA; WI; WV; WY; MB; NB; NS; ON; QC [Introduced in Europe]
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Discussion | Symphyotrichum cordifolium has been introduced in British Columbia but has not persisted. In a morphometric analysis of infraspecific variation in the northeastern part of the range, A. Legault (1986) showed that the varieties described are not distinct and mostly represent phenotypic variants caused by growing conditions; likewise, diploid and tetraploid races could not be distinguished morphologically. The type of Aster cordifolius var. laevigatus is conspecific with that of Symphyotrichum cordifolium. This name is the basionym of S. lowrieanum; therefore the latter cannot be considered distinct from S. cordifolium and is not recognized here. Most specimens initially identified as S. lowrieanum in herbaria have been re-determined as S. cordifolium. Some of the specimens, however, appear to correspond to the hybrid between S. cordifolium and S. laeve var. laeve, called S. ×schistosum (Steele) G. L. Nesom (syn. Aster schistosus Steele). Symphyotrichum ×tardiflorum (Linnaeus) Greuter, Aghababian & Wagenitz [syn. Aster tardiflorus Linnaeus, A. novi-belgii Linnaeus subsp. tardiflorus (Linnaeus) A. G. Jones, A. novi-belgii var. tardiflorus (Linnaeus) A. G. Jones, Symphyotrichum novi-belgii (Linnaeus) G. L. Nesom var. tardiflorum (Linnaeus) G. L. Nesom] is the F1 hybrid between S. cordifolium and S. puniceum (J. Labrecque & Brouillet 1996; G. L. Nesom 1997; W. Greuter et al. 2005). Aster tardiflorus forma vestitus (Fernald) Fernald or var. vestitus Fernald is a hairy variant of the hybrid and is not recognized here; hairiness may be related to whichever species is the maternal parent. A. G. Jones (1989) reported hybrids with S. drummondii and S. urophyllum. She believed that var. moratum may be a hybrid with or an introgressant from S. drummondii, but this is not accepted by all authors. Symphyotrichum ×finkii (Rydberg) G. L. Nesom (syn. Aster finkii Rydberg), may be the hybrid of S. cordifolium and S. shortii. All such putative hybrids need to be confirmed. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Symphyotrichum novae-angliae is escaped from cultivation and introduced in Montana, Oregon, Utah, Washington, and Wyoming, and has been reported as an ephemeral escape in British Columbia. It possibly escaped from cultivation elsewhere. The Michaelmas daisy is widely sold in the horticultural trade, where cultivars have been developed. Forms have been described that correspond to color genetic variants within natural populations {Aster novae-angliae forma roseus (Desfontaines) Britton; A. novae-angliae forma geneseensis House}; they are not recognized here. Symphyotrichum novae-angliae resembles Canadanthus modestus, but the ranges of the two do not overlap, and the latter has sparsely hairy cypselae with dark ribs. Symphyotrichum novae-angliae hybridizes with S. ericoides, forming the F1 intersectional hybrid S. ×amethystinum. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 20, p. 501. | FNA vol. 20, p. 487. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Aster cordifolius, Aster cordifolius var. alvearius, Aster cordifolius var. furbishiae, Aster cordifolius var. incisus, Aster cordifolius subsp. laevigatus, Aster cordifolius var. laevigatus, Aster cordifolius var. lanceolatus, Aster cordifolius var. moratus, Aster cordifolius var. polycephalus, Aster cordifolius var. racemiflorus, Aster cordifolius subsp. sagittifolius, Aster cordifolius var. sagittifolius, Aster finkii var. moratus, Aster leiophyllus, Aster lowrieanus, Aster lowrieanus var. incisus, Aster lowrieanus var. lanceolatus, Aster sagittifolius, S. cordifolium var. furbishiae, S. cordifolium var. lanceolatum, S. cordifolium var. moratum, S. cordifolium var. polycephalum, S. cordifolium var. racemiflorum, S. lowrieanum, S. sagittifolium | Aster novae-angliae, Virgulus novae-angliae |
Name authority | (Linnaeus) G. L. Nesom: Phytologia 77: 278. (1995) | (Linnaeus) G. L. Nesom: Phytologia 77: 287. (1995) |
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