Symphyotrichum cordifolium |
Symphyotrichum dumosum |
|
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aster cordifolié, common blue wood aster, heart-leaf American-aster, heart-leaf aster, heartleaf or common blue wood aster |
bushy American-aster, bushy aster, rice button aster |
|
Habit | Perennials, 20–120 cm, colonial or cespitose; branched rhizomatous, or with branched caudices, becoming ± woody. | Perennials, (20–)30–100 cm, colonial or cespitose; usually long-rhizomatous, often thick, woody, sometimes short, or with short, stout caudices. |
Stems | 1–5+, erect (straight to ± flexuous distally, often reddish, sometimes brown), usually glabrous, sometimes ± pilose, particularly distally. |
1–5+, erect (straight, often slender, brittle), densely or sparsely strigose to glabrescent or glabrate. |
Leaves | thin, margins serrate (often sharply, teeth acuminate, mucronulate) to serrate-crenate or subentire, strigose, apices acuminate to acute, mucronulate, abaxial faces glabrous or sparsely to densely strigose-pilose, often pilose on midveins, sometimes on other veins also, adaxial glabrous or sparsely to densely strigose, sometimes ± scabrous; basal withering by flowering, new vernal rosettes often present, long-petiolate (petioles ± narrowly winged, bases dilated, sheathing, ciliate), blades ovate to elliptic or suborbiculate, (10–)35–150 × (10–)25–75 mm, bases usually deeply cordate, sometimes rounded, margins coarsely, often irregularly serrate, apices sometimes obtuse or rounded; proximal cauline often withering by flowering, winged-petiolate (becoming shorter and more widely winged distally, petiole bases clasping), blades widely to narrowly ovate, 40–100(–140) × 20–40(–70) mm, reduced distally, bases ± deeply cordate to rounded, margins sharply serrate, apices acuminate; distal usually sessile or subsessile, rarely short-petiolate, blades ovate to lanceolate, 5–105 × 2–45 mm, bases cordate or rounded to attenuate or cuneate, margins serrate or entire (distalmost), apices acuminate. |
firm, margins entire to crenulate-serrate, recurved, scabrous, apices sharply white-mucronate, abaxial faces glabrous or glabrate to sparsely strigillose, adaxial glabrous or glabrate to submarginally scabrous (short-strigose), cauline withering by flowering (except on array branches), often with axillary leaf clusters; basal withering by flowering, petiolate, petioles winged, sheathing, strigose-ciliate, blades oblanceolate to spatulate (often declined), 8–50 × 3–15 mm, bases attenuate to cuneate, margins crenate-serrate, apices obtuse or rounded to ± acute; proximal cauline (mostly declined) subpetiolate (petioles widely winged, clasping) or sessile (then not or barely clasping), blades linear-oblanceolate, 25–120 × 1.5–9 mm, bases cuneate to slightly attenuate; distal (ascending or spreading) sessile, blades oblong or linear-lanceolate to linear, 2–45 × 0.5–4 mm, reduced distally (abruptly on branches), bases cuneate to rounded, not clasping, margins entire or serrate. |
Peduncles | 0.3–2 cm, ± pilose, bracts linear-oblanceolate or -lanceolate to linear, foliaceous, distally grading into phyllaries, margins sparsely ciliolate, glabrous. |
slender, usually stiff (sometimes lax), (0.5–)1–5 cm (rarely subsessile), progressively reduced distally, usually not secund, sparsely strigillose or glabrous, bracts 5–16+, spreading to ascending, linear-oblong or -elliptic to subulate or linear, progressively reduced distally, grading into phyllaries. |
Involucres | cylindro-campanulate to cylindric, (3–)4.5–5(–6) mm. |
cylindro-campanulate, (3–)4.5–6.3 mm. |
Ray florets | (8–)10–16(–20); corollas usually blue to purple, seldom whitish or pink, laminae (5–)6–8(–10) × 1.4–1.8 mm. |
15–33; corollas pale blue, pink, or lavender to white, laminae (4–)5–7(–8) × 1–1.7 mm. |
Disc florets | (8–)10–15(–20); corollas cream-color or light yellow becoming purple, (3–)4–4.5(–5) mm, tubes slightly shorter than funnelform throats, lobes sometimes ± spreading, narrowly triangular to lanceolate, 0.6–0.9 mm. |
15–30; corollas cream to pale yellow turning pink, 3.5–4.5 mm, tubes shorter than to equal to narrowly funnelform throats, lobes lanceolate, (0.4–)0.6–1.1 mm. |
Phyllaries | in (3–)4–6 series, linear-lanceolate to oblong-lanceolate, unequal, bases indurate 1/2–3/4, margins scarious, erose, hyaline, sparsely ciliolate, green zones lanceolate to ± diamond-shaped, apical, apices (often red-tipped) acute to obtuse-acuminate or acuminate, mucronulate, faces glabrous or sparsely strigillose. |
in 4–6 series, appressed or slightly spreading, oblong-oblanceolate (outer) to linear-oblanceolate or linear (innermost), strongly unequal, bases indurate (1/2–)2/3–4/5, margins hyaline, scarious, erose, distally ciliolate, green zones oblanceolate to elliptic (subapical), apices acute to obtuse (scarious), sometimes faintly reddish, mucronulate, faces usually glabrous, sometimes glabrate. |
Heads | [(5–)20–300+] in ± densely paniculiform arrays, branches divaricate to ascending, paniculiform, sometimes ± long-arching, leafy. |
in remote, diffuse, open, paniculiform arrays, branches numerous, ascending to stiffly ascending, secondary ones stiff, racemiform (usually not secund). |
Cypselae | dull purple or light brown, obovoid, ± compressed, 2–2.5 mm, 4–5-nerved, faces glabrous; pappi white or ± rose-tinged, 2.5–4.5 mm. |
pink or stramineous with pink streaks, or gray (± dark, nerves stramineous), oblong-obovoid, sometimes ± compressed, 1.5–2.5 mm, 3–4-nerved (nerves prominent), faces strigillose; pappi white to sordid, 4 mm. |
2n | = 16, 32. |
= 16, 32. |
Symphyotrichum cordifolium |
Symphyotrichum dumosum |
|
Phenology | Flowering Aug–Oct. | Flowering Aug–Oct. |
Habitat | Rich, mostly mesic, rocky to loamy soils, open wooded slopes and bluffs, stream banks, moist ledges, swampy woods, border of beech-maple or oak-hickory forets, clearings, thickets, roadsides, along ditches, sometimes weedy in urban areas | Open or wooded, moist or wet soils, bogs, fens, sedge meadows, marshes, swamps, flood plains, sandy or calcareous flats, loamy prairies, old fields, sandhills, flatwoods, hammocks, pine-hickory woods, oak or pine thickets, secondary woods, sandy to mucky or marly shores of lakes and ponds, interdunal hollows |
Elevation | 0–1200 m (0–3900 ft) | 0–700 m (0–2300 ft) |
Distribution |
AL; AR; CT; DC; GA; IA; IL; IN; KY; MA; MD; ME; MI; MN; MO; NC; NE; NH; NJ; NY; OH; OK; PA; RI; SC; SD; TN; VA; VT; WI; WV; MB; NB; NS; ON; PE; QC
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AL; AR; CT; DC; DE; FL; GA; IA; IL; IN; KY; LA; MA; MD; ME; MI; MO; MS; NC; NH; NJ; NY; OH; OK; PA; RI; SC; TN; TX; VA; WI; WV; NB; ON
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Discussion | Symphyotrichum cordifolium has been introduced in British Columbia but has not persisted. In a morphometric analysis of infraspecific variation in the northeastern part of the range, A. Legault (1986) showed that the varieties described are not distinct and mostly represent phenotypic variants caused by growing conditions; likewise, diploid and tetraploid races could not be distinguished morphologically. The type of Aster cordifolius var. laevigatus is conspecific with that of Symphyotrichum cordifolium. This name is the basionym of S. lowrieanum; therefore the latter cannot be considered distinct from S. cordifolium and is not recognized here. Most specimens initially identified as S. lowrieanum in herbaria have been re-determined as S. cordifolium. Some of the specimens, however, appear to correspond to the hybrid between S. cordifolium and S. laeve var. laeve, called S. ×schistosum (Steele) G. L. Nesom (syn. Aster schistosus Steele). Symphyotrichum ×tardiflorum (Linnaeus) Greuter, Aghababian & Wagenitz [syn. Aster tardiflorus Linnaeus, A. novi-belgii Linnaeus subsp. tardiflorus (Linnaeus) A. G. Jones, A. novi-belgii var. tardiflorus (Linnaeus) A. G. Jones, Symphyotrichum novi-belgii (Linnaeus) G. L. Nesom var. tardiflorum (Linnaeus) G. L. Nesom] is the F1 hybrid between S. cordifolium and S. puniceum (J. Labrecque & Brouillet 1996; G. L. Nesom 1997; W. Greuter et al. 2005). Aster tardiflorus forma vestitus (Fernald) Fernald or var. vestitus Fernald is a hairy variant of the hybrid and is not recognized here; hairiness may be related to whichever species is the maternal parent. A. G. Jones (1989) reported hybrids with S. drummondii and S. urophyllum. She believed that var. moratum may be a hybrid with or an introgressant from S. drummondii, but this is not accepted by all authors. Symphyotrichum ×finkii (Rydberg) G. L. Nesom (syn. Aster finkii Rydberg), may be the hybrid of S. cordifolium and S. shortii. All such putative hybrids need to be confirmed. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Symphyotrichum dumosum is introduced in New Brunswick. It is widely cultivated. It can hybridize with S. racemosum and S. lanceolatum var. interior (A. G. Jones 1989). G. L. Nesom (1994b) and J. C. Semple et al. (2002) recognized several varieties within the complex: var. dumosum [syn. Aster coridifolius Michaux, A. dumosus Linnaeus var. coridifolius (Michaux) Torrey & A. Gray]; var. gracilipes (Wiegand) G. L. Nesom (syn. A. dumosus var. gracilipes Wiegand); var. pergracile (Wiegand) G. L. Nesom (syn. A. dumosus var. pergracile Wiegand); var. strictior (Torrey & A. Gray) G. L. Nesom [syn. A. dumosus var. strictior Torrey & A. Gray, A. dumosus var. dodgei Fernald, S. dumosum var. dodgei (Fernald) G. L. Nesom, the latter possibly the hybrid S. dumosum × S. boreale fide G. L. Nesom 1997]; and var. subulifolium (Torrey & A. Gray) G. L. Nesom (syn. A. dumosus var. subulifolium Torrey & A. Gray). Some of these taxa may deserve higher rank. More work is needed in the complex, however, before a coherent taxonomy can be achieved. G. L. Nesom (1997) recognized also a southern pine flatwoods entity called Symphyotrichum kralii G. L. Nesom, based on the illegitimately named Aster pinifolius Alexander (not Nees, nor F. Mueller). The type of A. pinifolius, however, appears to be conspecific with that of S. simmondsii. At present, it seems best to wait for full investigation before formally recognizing such taxa. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 20, p. 501. | FNA vol. 20, p. 514. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Aster cordifolius, Aster cordifolius var. alvearius, Aster cordifolius var. furbishiae, Aster cordifolius var. incisus, Aster cordifolius subsp. laevigatus, Aster cordifolius var. laevigatus, Aster cordifolius var. lanceolatus, Aster cordifolius var. moratus, Aster cordifolius var. polycephalus, Aster cordifolius var. racemiflorus, Aster cordifolius subsp. sagittifolius, Aster cordifolius var. sagittifolius, Aster finkii var. moratus, Aster leiophyllus, Aster lowrieanus, Aster lowrieanus var. incisus, Aster lowrieanus var. lanceolatus, Aster sagittifolius, S. cordifolium var. furbishiae, S. cordifolium var. lanceolatum, S. cordifolium var. moratum, S. cordifolium var. polycephalum, S. cordifolium var. racemiflorum, S. lowrieanum, S. sagittifolium | Aster dumosus, Aster coridifolius |
Name authority | (Linnaeus) G. L. Nesom: Phytologia 77: 278. (1995) | (Linnaeus) G. L. Nesom: Phytologia 77: 280. (1995) |
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