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manyray aster

amethyst aster, aster, hybrid aster

Habit Perennials, 20–100(–120) cm, cespitose; usually with short, stout, branched caudices, woody with age, sometimes long-rhizomatous. Perennials, cespitose, 30–120 cm; with thick, woody, short-branched caudices at ends of fleshy rhizomes.
Stems

1–5+, erect or ascending (straight, often stout), proximally usually ± densely hirtellous, sometimes glabrescent, distally hirsute.

1–5+, erect (light brown to grayish brown), densely hispidulo-hirsute.

Leaves

thick, firm, margins (slightly undulate) scabrous, adaxial faces glabrous or strigose, scabrous, adaxial ± sparsely hirsute to scabro-hirtellous, sometimes ± scabrous (cauline sometimes with tufts of smaller leaves in axils);

basal withering by flowering, petiolate (petioles sometimes narrowly winged), blades oblong-ovate to lanceolate, 10–40 × 10–20 mm, bases deeply cordate, margins sparsely serrate to serrulate, apices usually acute, sometimes obtuse to rounded;

proximal cauline usually withering by flowering, petiolate (petioles slender to ± winged), blades ovate to broadly lanceolate, 40–90 × (15–) 25–50 mm, bases shallowly cordate or rounded to attenuate, rarely truncate, margins subentire or entire (rarely serrulate), apices acute or acuminate, with short callus point;

distal subpetiolate or sessile, blades ovate to lanceolate or linear-lanceolate, 10–70 × 1–10 mm, gradually or abruptly reduced distally, bases attenuate to cuneate, margins entire, apices acute to acuminate.

light green, thin to firm;

basal withering by flowering, sessile, blades oblanceolate, 10–40 × 3–10 mm, attenuate, margins usually entire, rarely remotely serrate, piloso-ciliate, apices obtuse, rounded to mucronulate, faces sparsely to moderately strigose;

proximal cauline sometimes persistent by flowering, sessile, blades linear or oblong to elliptic-oblanceolate, 40–60 × 3–10 mm, bases rounded or slightly clasping, margins entire, scabrous, apices acute to obtuse, faces ± copiously hirsute;

distal sessile, blades oblanceolate, 15–45 × 2–3(–4.5) mm, progressively reduced distally, bases cuneate, margins entire, apices acute, white-spine-tipped, faces sparsely strigose.

Peduncles

densely bracteate, 0.3–5(–10) cm (rarely subsesssile), bracts numerous, linear, 2–6 mm, mucronulate, grading into phyllaries.

hispid, bracts 1–3+, oblong-lanceolate, 3–5 mm, not grading into phyllaries.

Involucres

campanulate, 5–10 mm.

campanulate to hemispheric, 4–6 mm.

Ray florets

18–45;

corollas usually bright lavender-blue to purple, seldom white, laminae 9–15(–18) × 1–3.2 mm.

20–35;

corollas azure blue to violet or lavender, laminae 5–10 × 0.6–1.2 mm.

Disc florets

33–40+;

corollas cream or light yellow turning pinkish purple, (4–)4.5–5.5 mm, tubes shorter than funnelform throats, lobes sometimes ± spreading, triangular to lanceolate, 0.5–0.8 mm.

20–30+;

corollas light yellow turning pinkish to purple, 3–4 mm, tubes ca. 1/2 narrowly funnelform throats, lobes triangular, 0.5–0.7 mm, glabrous.

Phyllaries

in 4–6 series, oblong-lanceolate (outer) to oblanceolate-linear (innermost), unequal, bases indurate (appressed), margins ciliate to scabro-ciliate, green zones diamond-shaped to lanceolate, ± foliaceous distally, apices reflexed to squarrose, long-acuminate, apiculate, often purple, faces strigoso-hirtellous.

in 3–5 series, linear-lanceolate, ± unequal to subequal, bases ± indurate, margins hyaline proximally, outer distally hispido-scabrous, green zones diamond-shaped, restricted to distal 1/2–1/4, apices (outer) spreading to reflexed, acute, subspinulose, (inner) acuminate, purplish, faces (outer) scabroso-puberulent or strigose, eglandular, inner distally stipitate-glandular.

Heads

in open, diffuse, paniculiform arrays, branches divaricate, stout, long, densely bracteate.

in ample, leafy, paniculiform arrays, branches ascending, sometimes secund (heads often crowded).

Cypselae

deep purple to purplish brown or brown with purple speckles (nerves stramineous), oblong-obovoid, compressed, (1–)2.5–3.5(–3.8) mm, 5–6-nerved, faces glabrous;

pappi tawny or rose-tinged, 3.7–5.2 mm.

dull purple or brown, obovoid, not compressed, 1.5–2 mm, 7–9-nerved, faces densely sericeous;

pappi tan to tawny, sometimes rose to violet-tinged, 3.5–5.5 mm.

2n

= 16.

= 10.

Symphyotrichum anomalum

Symphyotrichum ×amethystinum

Phenology Flowering late Aug–Oct. Flowering Aug–Oct.
Habitat Dry, sandy, loamy, or clayey soils over limestone, or acid soils over chert, sandstone, or granite, rocky, open deciduous woods, thickets, dry ridges, cliffs, bluffs, occasional along streams Fields, prairies, disturbed grounds
Elevation 50–500 m (200–1600 ft) 200–400 m (700–1300 ft)
Distribution
from FNA
AR; IL; KS; MO; OK
[WildflowerSearch map]
[BONAP county map]
from FNA
CO; CT; IA; IL; IN; KY; MA; MD; MN; MO; ND; NE; NJ; NY; PA; RI; VT; WA; WI; ON
[BONAP county map]
Discussion

A. G. Jones (1989, 1992) reported that Symphyotrichum anomalum may hybridize with S. drummondii, S. oolentangiense, and S. shortii.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Symphyotrichum ×amethystinum is the F1 hybrid between S. ericoides and S. novae-angliae, encountered sometimes throughout the area where the two parental species co-occur. It is morphologically intermediate; it has non-spiny, sparsely stipitate-glandular phyllaries and rose-violet rays in mid-sized heads. Forma leucerythros Bemis and forma leucos Bemis have been described within this hybrid and may represent recombinants or normal population color variants.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 20, p. 504. FNA vol. 20, p. 497.
Parent taxa Asteraceae > tribe Astereae > Symphyotrichum > subg. Symphyotrichum > sect. Symphyotrichum Asteraceae > tribe Astereae > Symphyotrichum > subg. Virgulus
Sibling taxa
S. adnatum, S. anticostense, S. ascendens, S. boreale, S. bracteolatum, S. campestre, S. chapmanii, S. chilense, S. ciliatum, S. ciliolatum, S. concolor, S. cordifolium, S. cusickii, S. defoliatum, S. depauperatum, S. drummondii, S. dumosum, S. elliottii, S. ericoides, S. eulae, S. falcatum, S. fendleri, S. firmum, S. foliaceum, S. fontinale, S. frondosum, S. georgianum, S. grandiflorum, S. greatae, S. hallii, S. hendersonii, S. jessicae, S. laeve, S. lanceolatum, S. lateriflorum, S. laurentianum, S. lentum, S. molle, S. nahanniense, S. novae-angliae, S. novi-belgii, S. oblongifolium, S. ontarionis, S. oolentangiense, S. parviceps, S. patens, S. phlogifolium, S. pilosum, S. plumosum, S. porteri, S. potosinum, S. praealtum, S. pratense, S. prenanthoides, S. priceae, S. puniceum, S. pygmaeum, S. racemosum, S. retroflexum, S. rhiannon, S. robynsianum, S. sericeum, S. shortii, S. simmondsii, S. spathulatum, S. subspicatum, S. subulatum, S. tenuifolium, S. tradescantii, S. turbinellum, S. undulatum, S. urophyllum, S. walteri, S. welshii, S. yukonense, S. ×amethystinum
S. adnatum, S. anomalum, S. anticostense, S. ascendens, S. boreale, S. bracteolatum, S. campestre, S. chapmanii, S. chilense, S. ciliatum, S. ciliolatum, S. concolor, S. cordifolium, S. cusickii, S. defoliatum, S. depauperatum, S. drummondii, S. dumosum, S. elliottii, S. ericoides, S. eulae, S. falcatum, S. fendleri, S. firmum, S. foliaceum, S. fontinale, S. frondosum, S. georgianum, S. grandiflorum, S. greatae, S. hallii, S. hendersonii, S. jessicae, S. laeve, S. lanceolatum, S. lateriflorum, S. laurentianum, S. lentum, S. molle, S. nahanniense, S. novae-angliae, S. novi-belgii, S. oblongifolium, S. ontarionis, S. oolentangiense, S. parviceps, S. patens, S. phlogifolium, S. pilosum, S. plumosum, S. porteri, S. potosinum, S. praealtum, S. pratense, S. prenanthoides, S. priceae, S. puniceum, S. pygmaeum, S. racemosum, S. retroflexum, S. rhiannon, S. robynsianum, S. sericeum, S. shortii, S. simmondsii, S. spathulatum, S. subspicatum, S. subulatum, S. tenuifolium, S. tradescantii, S. turbinellum, S. undulatum, S. urophyllum, S. walteri, S. welshii, S. yukonense
Synonyms Aster anomalus Aster amethystinus, Virgulus ×amethystinus
Name authority (Engelmann ex Torrey & A. Gray) G. L. Nesom: Phytologia 77: 275. (1995) (Nuttall) G. L. Nesom: Phytologia 77: 294. (1995)
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