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manyray aster

Welsh's aster

Habit Perennials, 20–100(–120) cm, cespitose; usually with short, stout, branched caudices, woody with age, sometimes long-rhizomatous. Perennials, 30–100 cm colonial; rhizomes shallow and long, often relatively thick (those of the season each producing a distal rosette near the parent plant).
Stems

1–5+, erect or ascending (straight, often stout), proximally usually ± densely hirtellous, sometimes glabrescent, distally hirsute.

1–2, ascending to erect (sometimes lax, sometimes reddish), glabrous or sparsely strigoso-villosulous distally (arrays).

Leaves

thick, firm, margins (slightly undulate) scabrous, adaxial faces glabrous or strigose, scabrous, adaxial ± sparsely hirsute to scabro-hirtellous, sometimes ± scabrous (cauline sometimes with tufts of smaller leaves in axils);

basal withering by flowering, petiolate (petioles sometimes narrowly winged), blades oblong-ovate to lanceolate, 10–40 × 10–20 mm, bases deeply cordate, margins sparsely serrate to serrulate, apices usually acute, sometimes obtuse to rounded;

proximal cauline usually withering by flowering, petiolate (petioles slender to ± winged), blades ovate to broadly lanceolate, 40–90 × (15–) 25–50 mm, bases shallowly cordate or rounded to attenuate, rarely truncate, margins subentire or entire (rarely serrulate), apices acute or acuminate, with short callus point;

distal subpetiolate or sessile, blades ovate to lanceolate or linear-lanceolate, 10–70 × 1–10 mm, gradually or abruptly reduced distally, bases attenuate to cuneate, margins entire, apices acute to acuminate.

(adaxially green, abaxially pale green, sometimes glaucous) thin (proximal) to stiff (distal), margins serrulate (proximal) or entire or nearly so, ± revolute, scabrous, apices mucronulate or sometimes callous-pointed, faces glabrous;

basal withering by flowering, petiolate (petioles narrowly winged, sheathing), blades oblanceolate to spatulate, 10–40+ × 4–9+ mm, bases attenuate, margins entire, apices obtuse;

proximal cauline withering by flowering, sessile or winged-petiolate, blades lanceolate to oblanceolate, 50–150 × 8–11 mm, bases slightly clasping, margins sparsely serrulate, apices acute;

distal sessile, blades linear to linear-lanceolate, 50–130 × 4–6 mm, gradually reduced distally, bases ± clasping, apices acute.

Peduncles

densely bracteate, 0.3–5(–10) cm (rarely subsesssile), bracts numerous, linear, 2–6 mm, mucronulate, grading into phyllaries.

0.5–2 cm, scabro-villosulous, bracts linear, scabro-ciliolate.

Involucres

campanulate, 5–10 mm.

cylindro-campanulate, 4–7 mm.

Ray florets

18–45;

corollas usually bright lavender-blue to purple, seldom white, laminae 9–15(–18) × 1–3.2 mm.

18–25;

corollas pink to white, laminae 9–12 × 0.8–1.5 mm.

Disc florets

33–40+;

corollas cream or light yellow turning pinkish purple, (4–)4.5–5.5 mm, tubes shorter than funnelform throats, lobes sometimes ± spreading, triangular to lanceolate, 0.5–0.8 mm.

23–60;

corollas pale yellow, 3.8–6.4 mm, tubes shorter than funnelform throats, lobes narrowly triangular, 0.4–0.8 mm.

Phyllaries

in 4–6 series, oblong-lanceolate (outer) to oblanceolate-linear (innermost), unequal, bases indurate (appressed), margins ciliate to scabro-ciliate, green zones diamond-shaped to lanceolate, ± foliaceous distally, apices reflexed to squarrose, long-acuminate, apiculate, often purple, faces strigoso-hirtellous.

in 3–4 series, lanceolate (outer) to linear (inner), ± unequal, bases indurate, margins hyaline, erose, distally scabro-ciliolate to ciliolate, green zones ± foliaceous (outer) or lanceolate, apices acute to acuminate, sometimes purplish, faces glabrous.

Heads

in open, diffuse, paniculiform arrays, branches divaricate, stout, long, densely bracteate.

in open, slender, leafy, paniculiform or racemiform arrays, branches ascending.

Cypselae

deep purple to purplish brown or brown with purple speckles (nerves stramineous), oblong-obovoid, compressed, (1–)2.5–3.5(–3.8) mm, 5–6-nerved, faces glabrous;

pappi tawny or rose-tinged, 3.7–5.2 mm.

pinkish tan (nerves stramineous), obovoid, ± compressed, 0.8–1.6 mm, 3–5-nerved, faces sparsely strigillose;

pappi white, 2.8–5.3 mm.

2n

= 16.

= 16.

Symphyotrichum anomalum

Symphyotrichum welshii

Phenology Flowering late Aug–Oct. Flowering Aug–Oct.
Habitat Dry, sandy, loamy, or clayey soils over limestone, or acid soils over chert, sandstone, or granite, rocky, open deciduous woods, thickets, dry ridges, cliffs, bluffs, occasional along streams Wet soils in dry areas, hanging gardens, seeps, wet ledges, stream banks
Elevation 50–500 m (200–1600 ft) 1300–2300 m (4300–7500 ft)
Distribution
from FNA
AR; IL; KS; MO; OK
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; ID; MT; UT; WY
[BONAP county map]
Discussion

A. G. Jones (1989, 1992) reported that Symphyotrichum anomalum may hybridize with S. drummondii, S. oolentangiense, and S. shortii.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 20, p. 504. FNA vol. 20, p. 523.
Parent taxa Asteraceae > tribe Astereae > Symphyotrichum > subg. Symphyotrichum > sect. Symphyotrichum Asteraceae > tribe Astereae > Symphyotrichum > subg. Symphyotrichum > sect. Symphyotrichum
Sibling taxa
S. adnatum, S. anticostense, S. ascendens, S. boreale, S. bracteolatum, S. campestre, S. chapmanii, S. chilense, S. ciliatum, S. ciliolatum, S. concolor, S. cordifolium, S. cusickii, S. defoliatum, S. depauperatum, S. drummondii, S. dumosum, S. elliottii, S. ericoides, S. eulae, S. falcatum, S. fendleri, S. firmum, S. foliaceum, S. fontinale, S. frondosum, S. georgianum, S. grandiflorum, S. greatae, S. hallii, S. hendersonii, S. jessicae, S. laeve, S. lanceolatum, S. lateriflorum, S. laurentianum, S. lentum, S. molle, S. nahanniense, S. novae-angliae, S. novi-belgii, S. oblongifolium, S. ontarionis, S. oolentangiense, S. parviceps, S. patens, S. phlogifolium, S. pilosum, S. plumosum, S. porteri, S. potosinum, S. praealtum, S. pratense, S. prenanthoides, S. priceae, S. puniceum, S. pygmaeum, S. racemosum, S. retroflexum, S. rhiannon, S. robynsianum, S. sericeum, S. shortii, S. simmondsii, S. spathulatum, S. subspicatum, S. subulatum, S. tenuifolium, S. tradescantii, S. turbinellum, S. undulatum, S. urophyllum, S. walteri, S. welshii, S. yukonense, S. ×amethystinum
S. adnatum, S. anomalum, S. anticostense, S. ascendens, S. boreale, S. bracteolatum, S. campestre, S. chapmanii, S. chilense, S. ciliatum, S. ciliolatum, S. concolor, S. cordifolium, S. cusickii, S. defoliatum, S. depauperatum, S. drummondii, S. dumosum, S. elliottii, S. ericoides, S. eulae, S. falcatum, S. fendleri, S. firmum, S. foliaceum, S. fontinale, S. frondosum, S. georgianum, S. grandiflorum, S. greatae, S. hallii, S. hendersonii, S. jessicae, S. laeve, S. lanceolatum, S. lateriflorum, S. laurentianum, S. lentum, S. molle, S. nahanniense, S. novae-angliae, S. novi-belgii, S. oblongifolium, S. ontarionis, S. oolentangiense, S. parviceps, S. patens, S. phlogifolium, S. pilosum, S. plumosum, S. porteri, S. potosinum, S. praealtum, S. pratense, S. prenanthoides, S. priceae, S. puniceum, S. pygmaeum, S. racemosum, S. retroflexum, S. rhiannon, S. robynsianum, S. sericeum, S. shortii, S. simmondsii, S. spathulatum, S. subspicatum, S. subulatum, S. tenuifolium, S. tradescantii, S. turbinellum, S. undulatum, S. urophyllum, S. walteri, S. yukonense, S. ×amethystinum
Synonyms Aster anomalus Aster welshii
Name authority (Engelmann ex Torrey & A. Gray) G. L. Nesom: Phytologia 77: 275. (1995) (Cronquist) G. L. Nesom: Phytologia 77: 294. (1995)
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